Aphodiinae
Aphodiinae is a subfamily of the scarab beetle family, Scarabaeidae. Members of this subfamily are known commonly as the small dung beetles and many, but not all, are dung beetles. These beetles are found worldwide.

Udo Schmidt from Deutschland · CC BY-SA 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ScarabaeidaePBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 121 million years on record.
The record covers at least 121.4 Mya to 129 ka. No occurrence read is dated outside 125.77 Mya to 11.7 ka.
182 occurrences of Aphodiinae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
182 occurrences- Early Cretaceous2%
- Paleoceneunder 1%
- Eocene20%
- Oligocene2%
- Miocene5%
- Plioceneunder 1%
- Pleistocene70%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
22 formations
- Florissant Formation25
- Highbury Silts and Sands Formation23
- Keremesit Formation8
- Olympia beds Formation7
- Titusville Till Formation5
- Eifel Formation4
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The named rock units Aphodiinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 87 occurrences have no formation on record, which is common for older literature.
Aphodiinae and its relatives
PleistoceneWhere Aphodiinae and its relatives were recorded in the Pleistocene.
Found in the same rock
3,617 taxa- Aphodius40 sites
- Morychus viridis21 sites
- Curtonotus alpinus19 sites
- Pterostichus (Cryobius)19 sites
- Carabidae18 sites
- Poecilus nearcticus18 sites
- Bembidion17 sites
- Coleoptera17 sites
- Phyllobius16 sites
- Stephanocleonus eruditus16 sites
- Chrysolina rufilabris15 sites
- Hypera ornata15 sites
- Staphylinidae15 sites
- Curculionidae13 sites
- Stephanocleonus foveifrons13 sites
- Cholevinus sibiricus12 sites
- Elateridae12 sites
- Coniocleonus ferrugineus11 sites
- Lepyrus nordenskioldi11 sites
- Stephanocleonus fossulatus11 sites
- Troglocollops arcticus11 sites
- Chrysomelidae10 sites
- Harpalus amputatus amputatoides10 sites
- Tachinus arcticus10 sites
The Paleobiology Database records 112 sites for Aphodiinae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
49 references- Late Cenozoic insects of northern Eurasia27
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK23
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian12
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA11
- Environmental interpretation of fossil insect assemblages from MIS 5 at Ziegler Reservoir, Snowmass Village, Colorado9
- New Miocene Coleoptera from Florissant9
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.7
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire7
- Paleogeography of the lower Volga at the end of Atelian time based on entomological and botanical data6
- Pleistoceńskie owady z Borysławia6
- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania5
- Fossil Coleoptera from the Wilson Ranch near Florissant, Colorado5
Each of the 182 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 37 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Aphodiinae
- Source
- Wikipedia/Commons
- Credit
- Udo Schmidt from Deutschland
- Licence
- CC BY-SA 2.0
- Commercial use
- the licence allows it
No 3D scan of Aphodiinae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Aphodiinae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itScarabaeidae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Leach 1815