Archipolypoda
Archipolypoda is an extinct group of millipedes known from fossils in Europe and North America and containing the earliest known land animals. The Archipolypoda was erected by Scudder (1882) but redefined in 2005 with the description of several new species from Scotland. Distinguishing characteristics include relatively large eyes with densely packed ocelli, and modified leg pairs on the 8th body ring. Some species had prominent spines while others had a flattened appearance.

John Sterling Kingsley · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- HelminthomorphaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
Time the record covers
At least 116 million years on record.
The record covers at least 426.7 Mya to 309.8 Mya. No occurrence read is dated outside 432.9 Mya to 307 Mya.
105 occurrences of Archipolypoda on record, most of them in the Carboniferous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
105 occurrences- Silurian9%
- Devonian11%
- Carboniferous80%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
13 formations
- Carbondale Formation · Kewanee Group43
- Kladno Formation16
- Cowie Formation · Stonehaven Group8
- Dundee Flagstone Formation · Arbuthnott-Garvock Group6
- Ballagan Formation5
- Pennant Formation2
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The named rock units Archipolypoda has been recovered from, largest first. Drawn where the rock is now, not where it formed. 17 occurrences have no formation on record, which is common for older literature.
Archipolypoda and its relatives
CarboniferousWhere Archipolypoda and its relatives were recorded in the Carboniferous.
Found in the same rock
360 taxa- Euphoberia armigera9 sites
- Acantherpestes major6 sites
- Architarbus rotundatus5 sites
- Euphoberia ferox5 sites
- Euphoberia granosa5 sites
- Kampecaris forfarensis5 sites
- Pleurojulus biornatus5 sites
- Eucaenus ovalis4 sites
- Palaeocampa anthrax4 sites
- Adiphlebia lacoana3 sites
- Albadesmus almondi3 sites
- Aphantomartus pustulatus3 sites
- Archidesmus macnicoli3 sites
- Climatius reticulatus3 sites
- Eoscorpius carbonarius3 sites
- Euphoberia3 sites
- Euphoberia anguilla3 sites
- Euphoberia carri3 sites
- Euproops danae3 sites
- Ischnacanthus gracilis3 sites
- Mesacanthus mitchelli3 sites
- Miamia bronsoni3 sites
- Palaeocaris typus3 sites
- Palaeosoma robustum3 sites
The Paleobiology Database records 51 sites for Archipolypoda, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
27 references- New Carboniferous Myriapoda from Illinois21
- Archipolypoda, a subordinal type of spined myriapods from the Carboniferous formation17
- Fritsch, 189915
- Morphology and taxonomy of Paleozoic millipedes (Diplopoda: Chilognatha: Archipolypoda) from Scotland8
- A new terrestrial millipede fauna of earliest Carboniferous (Tournaisian) age from southeastern Scotland helps fill ‘Romer's Gap'5
- Taxonomy and trunk-ring architecture of pleurojulid millipedes (Diplopoda: Chilognatha: Pleurojulida) from the Pennsylvanian of Europe and North America5
- On some fossil myriapods from the Lower Old Red Sandstone of Forfarshire4
- Age of the basal ‘Lower Old Red Sandstone’ Stonehaven Group of Scotland: The oldest reported air-breathing land animal is Silurian (late Wenlock) in age3
- Fossil myriopods from the Middle Coal-Measures of Spath Bottoms, Rochdale, Lancashire3
- A new genus of archipolypodan millipede from the Coseley Lagerstätte, Upper Carboniferous, UK2
- New flat-backed archipolypodan millipedes from the Upper Devonian of North America2
- The Silurian-Devonian fossil record of the Myriapoda2
Each of the 105 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 15 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Archipolypoda
- Source
- Wikipedia/Commons
- Credit
- John Sterling Kingsley
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Archipolypoda was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itHelminthomorpha through timeFormations of the CarboniferousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Scudder 1882)