Archostemata
The Archostemata are the smallest suborder of beetles, consisting of 50 living species in five families and over 200 described fossil species. They are an ancient lineage with a number of primitive characteristics. Antennae may be thread-shaped (filiform) or like a string of beads (moniliform). This suborder also contains the only beetles where both sexes are paedogenic, Micromalthus debilis. Modern archostematan beetles are considered rare, but were more diverse during the Mesozoic.

me · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- MetacoleopteraPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 289 million years on record.
The record covers at least 293.52 Mya to 3.6 Mya. No occurrence read is dated outside 298.9 Mya to 2.58 Mya.
731 occurrences of Archostemata on record, most of them in the Early Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
731 occurrences- Permian12%
- Early Triassic2%
- Middle Triassic10%
- Late Triassic8%
- Early Jurassic6%
- Middle Jurassic4%
- Late Jurassic14%
- Early Cretaceous27%
- Late Cretaceous8%
- Paleocene3%
- Eocene5%
- Oligoceneunder 1%
- Miocene1%
- Plioceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
119 formations
- Madygen Formation57
- Karabastau Formation46
- Dzun-Bain Formation · Bon Tsaagan Group40
- Yixian Formation · Jehol Group38
- Daohugou Formation25
- Menat Formation22
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The named rock units Archostemata has been recovered from, largest first. Drawn where the rock is now, not where it formed. 95 further formations hold fewer occurrences each and are not listed. 116 occurrences have no formation on record, which is common for older literature.
Archostemata and its relatives
Early CretaceousWhere Archostemata and its relatives were recorded in the Early Cretaceous.
Found in the same rock
7,297 taxa- Coleoptera17 sites
- Blattodea16 sites
- Insecta10 sites
- Cupedidae9 sites
- Notocupes elegans9 sites
- Odonata9 sites
- Orthoptera9 sites
- Aktassoblatta fusca8 sites
- Notocupes8 sites
- Blattula brevicaudata7 sites
- Procramptonomyia kovalevi7 sites
- Aeschnogomphus intermedius6 sites
- Allophalerus bontsaganensis6 sites
- Chresmoda obscura6 sites
- Cymatophlebia longialata6 sites
- Ditomoptera dubia6 sites
- Hexagenites cellulosus6 sites
- Isophlebia aspasia6 sites
- Lithoblatta lithophila6 sites
- Mesonepa primordialis6 sites
- Myrmicium schroeteri6 sites
- Nuurcala popovi6 sites
- Omma liassicum6 sites
- Pseudohydrophilus avitus6 sites
The Paleobiology Database records 289 sites for Archostemata, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
185 references- Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]69
- New Mesozoic cupedid beetles from Mongolia: Brochocoleini and Notocupedini36
- Paleozoiskie nasekomye kuznetskogo basseina36
- New beetles of the family Cupedidae from the Mesozoic of Mongoloa. Ommatini, Mesocupedini, Priacmini29
- Zhuki-Arkhostematy Yury Karatau (Coleoptera, Archostemata) [Jurassic Archostematid beetles from Karatau (Coleoptera, Archostemata)]27
- New beetles (Insecta, Coleoptera) from the latter half of the Permian of European Russia22
- Taxonomy of the reticulate beetles of the subfamily Cupedinae (Coleoptera: Archostemata), with a review of the historical development21
- A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata)16
- Noviye zhuki semeistva Cupedidae iz Yurskikh otlozheniy Karatau16
- Mesozoic insects of Queensland Part I - introduction and Coleoptera14
- Upper Jurassic Lagerstätte Shar Teg, southwestern Mongolia13
- New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family13
Each of the 731 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 173 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Archostemata
- Source
- Wikipedia/Commons
- Credit
- me
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Archostemata was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMetacoleoptera through timeFormations of the Early CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Kolbe 1908