Borophagus secundus
Borophagus secundus is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America during the Late Miocene to Early Pliocene from 9 to 5 Ma. Borophagus secundus existed for approximately 4 million years.

Xiaoming Wang Is a corresponding author, Stuart C White, Mairin Balisi, Jacob Biewer, Julia Sankey, Dennis Garber, Z… · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- BorophagusPBDB
- Diet
- carnivorePBDB, for Borophagus
- Life habit
- ground dwellingPBDB, for Canidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At most 7.8 million years on record.
Every occurrence read is dated to somewhere between 12.5 Mya and 4.7 Mya, and nothing in the dating separates them, so that window is what the record shows rather than a span.
35 occurrences of Borophagus secundus on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
35 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
13 formations
- Gracias Formation4
- Ogallala Formation3
- Rancho Viejo Beds Formation3
- Snake Creek Formation2
- Goodnight Beds Formation1
- Punchbowl Formation1
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The named rock units Borophagus secundus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 14 occurrences have no formation on record, which is common for older literature.
Borophagus secundus and its relatives
MioceneWhere Borophagus secundus and its relatives were recorded in the Miocene.
Found in the same rock
242 taxa- Pleiolama vera12 sites
- Neohipparion eurystyle11 sites
- Agriotherium schneideri9 sites
- Megatylopus matthewi9 sites
- Astrohippus stockii8 sites
- Dinohippus mexicanus8 sites
- Eucyon ferox8 sites
- Amphimachairodus coloradensis7 sites
- Teleoceras hicksi7 sites
- Alforjas6 sites
- Desmathyus brachydontus6 sites
- Dinohippus leidyanus6 sites
- Eucyon davisi6 sites
- Plesiogulo marshalli6 sites
- Tayassuinae6 sites
- Teleoceras fossiger6 sites
- Vulpes stenognathus6 sites
- Aphelops mutilus5 sites
- Astrohippus ansae5 sites
- Dinohippus interpolatus5 sites
- Hexobelomeryx fricki5 sites
- Mylagaulidae5 sites
- Nannippus aztecus5 sites
- Pediomeryx hemphillensis5 sites
The Paleobiology Database records 34 sites for Borophagus secundus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
13 references- Synonymies and reidentifications of North American fossil mammals15
- Phylogenetic systematics of the Borophaginae (Carnivora: Canidae)5
- The Early-Late Hemphillian (Hh2) Faunal Assemblage From Juchipila Basin, State of Zacatecas, Mexico, and Its Biochronologic Correlation with Other Hemphillian Faunas in Central Mexico3
- Increased xenarthran diversity of the Great American Biotic Interchange: a new genus and species of ground sloth (Mammalia, Xenarthra, Megalonychidae) from the Hemphillian (late Miocene) of Jalisco, Mexico2
- Late Cenozoic Vertebrates from Honduras and El Salvador2
- New occurrences of mammals from McKay Reservoir (Hemphillian, Oregon)1
- Contribución al conocimiento de los rinocerontes fósiles de la Cuenca de Tecolotlán, en el estado de Jalisco, México1
- Miocene Vertebrates From Ogallala Formation Sites in Western Oklahoma1
- Late Tertiary Canids from Central Mexico1
- Stratigraphy and biostratigraphy of late Cenozoic deposits in central Sioux County, western Nebraska1
- Mammalian fauna of the Drewsey Formation, Bartlett Mountain, Drinkwater and Otis Basin local faunas1
- Notes on Additional Fauna of Edson Quarry of the Middle Pliocene of Kansas1
Each of the 35 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 1 further reference is not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Borophagus secundus
- Source
- Wikipedia/Commons
- Credit
- Xiaoming Wang Is a corresponding author, Stuart C White, Mairin Balisi, Jacob Biewer, Julia Sankey, Dennis Garber, Z…
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
Laser scan
- Source
- MorphoSource
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Borophagus secundus · YPM:VP:10580
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itBorophagus through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (VanderHoof 1931)