Brontotheriidae
Brontotheres, also called titanotheres, are an extinct group of perissodactyl mammals belonging to the family Brontotheriidae. Brontotheres are traditionally interpreted as relatives of horses, though high-level perissodactyl relationships are not well understood. Brontotheres lived during the Eocene epoch and their fossils have mainly been found in Asia and North America, with some fossils also found in Eastern Europe. Early brontotheres were small animals, but later brontotheres grew to become the largest land mammals of the Eocene, and the largest land mammals to have evolved up until their time.

Henry Fairfield Osborn (1857–1935) · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- brontotherePBDB
- Sits inside
- PerissodactylaPBDB
- Diet
- browserPBDB
- Life habit
- ground dwellingPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 23 million years on record.
The record covers at least 51.1 Mya to 27.3 Mya. No occurrence read is dated outside 56 Mya to 23.04 Mya.
578 occurrences of Brontotheriidae on record, most of them in the Eocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
578 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
91 formations
- Bridger Formation55
- Uinta Formation38
- Chadron Formation · White River Group34
- Wind River Formation33
- Wasatch Formation29
- White River Formation23
Loading coastlines…
The named rock units Brontotheriidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 67 further formations hold fewer occurrences each and are not listed. 50 occurrences have no formation on record, which is common for older literature.
Brontotheriidae and its relatives
EoceneWhere Brontotheriidae and its relatives were recorded in the Eocene.
Found in the same rock
969 taxa- Lambdotherium popoagicum61 sites
- Palaeosyops fontinalis35 sites
- Hyopsodus30 sites
- Hyracotherium vasacciense30 sites
- Hyopsodus paulus27 sites
- Microsyops scottianus26 sites
- Paramys copei25 sites
- Coryphodon24 sites
- Hyrachyus modestus21 sites
- Esthonyx acutidens20 sites
- Heptodon posticus19 sites
- Hyopsodus wortmani19 sites
- Hyrachyus eximius19 sites
- Heptodon calciculus18 sites
- Bunophorus sinclairi17 sites
- Didymictis altidens17 sites
- Hyopsodus minusculus16 sites
- Perissodactyla16 sites
- Viverravus gracilis16 sites
- Diacodexis secans15 sites
- Phenacodus15 sites
- Cantius nunienus14 sites
- Eotitanops borealis14 sites
- Microsyops elegans14 sites
The Paleobiology Database records 467 sites for Brontotheriidae, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
201 references- Synonymies and reidentifications of North American fossil mammals75
- New Eurasiatic and American proboscideans19
- Summary of known occurrences of terrestrial vertebrates from Eocene strata of southern California16
- Early Eocene Mammalian Fauna of the Piceance Creek Basin. Northwestern Colorado15
- The Eocene Vertebrates of the Uinta Basin, Utah14
- Mammalian Fauna From the Lower Bridger Formation (Bridger A, Early Middle Eocene) of the Southern Green River Basin, Wyoming13
- Revision of the Wind River faunas, early Eocene of central Wyoming. Part 5. Geology and biostratigraphy of the upper part of the Wind River Formation, northeastern Wind River Basin12
- Fossil Mammalia of the Huerfano Formation, Eocene, of Colorado12
- Paleontology and Correlation of the Eocene Tepee Trail and Wiggins Formations in the North Fork of Owl Creek Area, Southeastern Absaroka Range, Hot Springs County, Wyoming11
- Geology and mammalian paleontology of the New Fork-Big Sandy area, Sublette County, Wyoming10
- Brontotheriidae (Perissodactyla) from the late early and middle Eocene (Brigerian), Wasatch and Bridger formations, southern Green River Basin, southwestern Wyoming9
- Fossil mammals from Burma in the American Museum of Natural History9
Each of the 578 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 189 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Brontotheriidae
- Source
- Wikipedia/Commons
- Credit
- Henry Fairfield Osborn (1857–1935)
- Licence
- Public Domain
- Commercial use
- the licence allows it
3D model
- Source
- Sketchfab
- Licence
- CC Attribution-NonCommercial
- Commercial use
- the licence does not allow it
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPerissodactyla through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Marsh 1873