Galericinae
Gymnures, also called hairy hedgehogs or moonrats, are mammals belonging to the subfamily Galericinae, in the family Erinaceidae and the order Eulipotyphla. Gymnures may resemble rats but are not closely related; they are not rodents at all, but instead are closely related to hedgehogs, being their sister group within Erinaceidae. They are thought to have appeared in Eastern Asia before their closest relatives, and changed little from the original ancestor, which is thought to have been also the ancestor of the shrews.

Mark Bolnik · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ErinaceidaePBDB
- Diet
- insectivore, carnivorePBDB, for Erinaceidae
- Life habit
- semifossorialPBDB, for Erinaceidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 42 million years on record.
The record covers at least 48.07 Mya to 5.09 Mya. No occurrence read is dated outside 56 Mya to 11.7 ka.
200 occurrences of Galericinae on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
200 occurrences- Eocene7%
- Oligocene21%
- Miocene65%
- Pliocene6%
- Pleistoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
41 formations
- Brule Formation7
- facies à Congeria Formation7
- Wagon Bed ("Tepee Trail") Formation5
- Lefkon Formation3
- Huadian Formation3
- Renova Formation2
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The named rock units Galericinae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 17 further formations hold fewer occurrences each and are not listed. 134 occurrences have no formation on record, which is common for older literature.
Galericinae and its relatives
MioceneWhere Galericinae and its relatives were recorded in the Miocene.
Found in the same rock
1,918 taxa- Deinogalerix intermedius34 sites
- Apodemus26 sites
- Lagomorpha20 sites
- Mikrotia19 sites
- Solitudo19 sites
- Schizogalerix18 sites
- Soricidae18 sites
- Prolagus16 sites
- Anguinae13 sites
- Lacerta13 sites
- Chiroptera12 sites
- Deinogalerix minor12 sites
- Latonia12 sites
- Neocricetodon12 sites
- Spermophilinus bredai12 sites
- Apodemus gudrunae11 sites
- Glis minor11 sites
- Paraethomys11 sites
- Rhinocerotidae11 sites
- Colubrinae10 sites
- Erinaceinae10 sites
- Muscardinus vireti10 sites
- Ophisaurus10 sites
- Plesiodimylus chantrei10 sites
The Paleobiology Database records 174 sites for Galericinae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
88 references- The giant erinaceid insectivore, Deinogalerix Freudenthal, from the Upper Miocene of Gargano, Italy51
- Insectivora und Chiroptera (Mammalia) aus dem Neogen der Türkei10
- Le Garouillas et les sites contemporains (Oligocene, MP 25) des phosphorites du Quercy (Lot, Tarn-et garonne, France) et leurs faunes de vertebres. 4. Marsupiaux et insectivores.9
- Late Messinian mollusks and vertebrates from Moncucco Torinese, north-western Italy. Paleoecological and paleoclimatological implications7
- Erinaceidae and Dimylidae (Lipotyphla) from the Upper Middle Miocene of South Germany6
- Insectivores pliocènes du sud de la France (Languedoc-Roussillon) et du nord-est de l'Espagne.6
- Greece4
- Synonymies and reidentifications of North American fossil mammals3
- First Paleogene Mammalian Fauna from Northeast China3
- On the classification of early Tertiary Erinaceomorpha (Insectivora, Mammalia)3
- Casa del Acero: nueva fuana Turoliense de vertebrados (Mioceno Superior de Fortuna, Murcia)3
- Les Insectivores des Phosphorites du Quercy. [Insectivores of the Quercy Phosphorites].3
Each of the 200 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 76 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Galericinae
- Source
- Wikipedia/Commons
- Credit
- Mark Bolnik
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Galericinae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Gymnure”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itErinaceidae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Pomel 1848