Litopterna
Litopterna is an extinct order of South American native ungulates that lived from the Paleocene to the Pleistocene-Holocene around 62.5 million to 12,000 years ago, and were also present in West Antarctica during the Eocene. They represent the second most diverse group of South American ungulates after Notoungulata. It is divided into nine families, with Proterotheriidae and Macraucheniidae being the most diverse and last surviving families.

Ivan Iofrida · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PanameriungulataPBDB
- Diet
- herbivorePBDB
- Life habit
- scansorialPBDB, for Eutheria
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 61 million years on record.
The record covers at least 61.66 Mya to 11.7 ka. No occurrence read is dated outside 66 Mya to 0 Mya.
583 occurrences of Litopterna on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
583 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
107 formations
- Santa Cruz Formation84
- Sarmiento Formation42
- Cerro Azul Formation25
- La Victoria Formation · Honda Group22
- Pinturas Formation21
- Monte Hermoso Formation17
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The named rock units Litopterna has been recovered from, largest first. Drawn where the rock is now, not where it formed. 83 further formations hold fewer occurrences each and are not listed. 132 occurrences have no formation on record, which is common for older literature.
Litopterna and its relatives
MioceneWhere Litopterna and its relatives were recorded in the Miocene.
Found in the same rock
2,122 taxa- Neoreomys australis23 sites
- Macrauchenia patagonica22 sites
- Hapalops19 sites
- Macraucheniidae19 sites
- Proterotheriidae19 sites
- Protypotherium19 sites
- Nesodon17 sites
- Adinotherium16 sites
- Macrauchenia16 sites
- Promacrauchenia15 sites
- Protypotherium praerutilum15 sites
- Theosodon15 sites
- Acdestis oweni14 sites
- Hippidion14 sites
- Palaeothentes minutus14 sites
- Smilodon populator14 sites
- Toxodon14 sites
- Astrapotherium13 sites
- Cladosictis patagonica13 sites
- Megatherium americanum13 sites
- Prozaedyus13 sites
- Testudines13 sites
- Thoatherium minusculum13 sites
- Acarechimys minutissimus12 sites
The Paleobiology Database records 295 sites for Litopterna, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
210 references- The beginning of the age of mammals in South America. Part I31
- Paleobiology of Santacrucian native ungulates (Meridiungulata: Astrapotheria, Litoptera and Notoungulata)25
- Los Litopterna (Mammalia) de la Formación Pinturas, Mioceno temprano-medio de Patagonia21
- Bioestratigrafía de la Formación Santa Cruz (Mioceno Inferior) en el extremo sudeste de la Patagonia20
- Updated data on Litopterna from the Huayquerian Stage/Age (Late Miocene-Early Pliocene) of central-east Argentina18
- Litopterns16
- Litopterna (Mammalia) From the Santa Cruz Formation (Early-Middle Miocene) At the Río Santa Cruz, Southern Argentina14
- Proterotheriidae (Mammalia, Litopterna) from the Cerro Azul Formation (late Miocene), La Pampa Province, Argentina12
- Miocene vertebrates from Entre Rios province, eastern Argentina12
- The Eocene to Pleistocene vertebrates of Bolivia and their stratigraphic context: a review12
- Biostratigraphy and biochronology of the Monte Hermoso Formation (early Pliocene) at its type locality, Buenos Aires Province, Argentina10
- Systematics of the Adianthidae (Litopterna, Mammalia)10
Each of the 583 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 198 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Litopterna
- Source
- Wikipedia/Commons
- Credit
- Ivan Iofrida
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Litopterna was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Litopterna”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPanameriungulata through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Ameghino 1889)