Meganisoptera
Meganisoptera is an extinct order of large dragonfly-like insects, informally known as griffenflies, griffinflies, or (incorrectly) as giant dragonflies. The order was formerly named Protodonata, the "proto-Odonata", for their similar appearance and supposed relation to modern Odonata. They range in Palaeozoic times. Though most were only slightly larger than modern dragonflies, the order includes the largest known insect species, such as the late Carboniferous Meganeura monyi and the even larger early Permian Meganeuropsis permiana, with wingspans of up to 71 centimetres (28 in).

André Nel, Jakub Prokop, Martina Pecharová, Michael S. Engel & Romain Garrouste · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- griffinflyPBDB
- Sits inside
- NeodonatopteraPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 55 million years on record.
The record covers at least 320 Mya to 264.28 Mya. No occurrence read is dated outside 323.4 Mya to 259.857 Mya.
97 occurrences of Meganisoptera on record, most of them in the Permian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
97 occurrences- Carboniferous44%
- Permian56%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
26 formations
- Wellington Formation · Cimarron Group27
- Salagou Formation12
- Commentry Shales Formation11
- Carbondale Formation · Kewanee Group5
- Yanghugou Formation4
- Iva-Gora Beds Formation4
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The named rock units Meganisoptera has been recovered from, largest first. Drawn where the rock is now, not where it formed. 2 further formations hold fewer occurrences each and are not listed. 10 occurrences have no formation on record, which is common for older literature.
Meganisoptera and its relatives
PermianWhere Meganisoptera and its relatives were recorded in the Permian.
Found in the same rock
1,090 taxa- Artinska clara6 sites
- Artinska ovata6 sites
- Dichentomum tinctum6 sites
- Lemmatophora typa6 sites
- Lisca minuta6 sites
- Asthenohymen dunbari5 sites
- Delopsocus elongatus5 sites
- Delopterum minutum5 sites
- Kennedya mirabilis5 sites
- Lecorium elongatum5 sites
- Liomopterum ornatum5 sites
- Parablattelytron rectum5 sites
- Permopsocus latipennis5 sites
- Permopsylla americana5 sites
- Protohymen permianus5 sites
- Tupus permianus5 sites
- Asthenohymen affinis4 sites
- Asthenohymen pusillus4 sites
- Blattodea4 sites
- Elmoa trisecta4 sites
- Misthodotes obtusus4 sites
- Oligotypus tillyardi4 sites
- Parablattelytron subincisum4 sites
- Paraprisca fragilis4 sites
The Paleobiology Database records 62 sites for Meganisoptera, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
46 references- Revision of Permo-Carboniferous griffenflies (Insecta: Odonatoptera: Meganisoptera) based upon new species and redescription of selected poorly known taxa from Eurasia19
- Revision der Paläozoischen Insekten7
- Lower Permian Insects from Oklahoma. Part 1. Introduction and the Orders Megasecoptera, Protodonata, and Odonata5
- The Lower Permian Insects of Kansas. Part 6: Delopteridae, Protelytroptera, Plectoptera and a New Collection of Protodonata, Odonata, Megasecoptera, Homoptera and Psocoptera5
- Kansas Permian Insects, Part 5: The Orders Protodonata and Odonata5
- The Lower Permian Insects of Kansas. Part 2: The Orders Paleodictyoptera, Protodonata and Odonata4
- Neue Meganeuridae aus dem Namurium von Hagen-Vorhalle (BRD) und die Phylogenie der Meganisoptera3
- New representatives of fossil insects of order Protodonata3
- Notes on the Protodonata and Protozygoptera of the Lower Permian of Kansas3
- The Lower Permian Insects of Kansas. Part 8: Additional Megasecoptera, Protodonata, Odonata, Homoptera, Psocoptera, Protelytroptera, Plectoptera and Protoperlaria3
- Early Pennsylvanian Odonatoptera from the Xiaheyan locality (Ningxia, China): new material, taxa, and perspectives2
- Carboniferous protodonatoid dragonfly nymphs and the synapomorphies of Odonatoptera and Ephemeroptera (Insecta: Palaeoptera)2
Each of the 97 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 34 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Meganisoptera
- Source
- Wikipedia/Commons
- Credit
- André Nel, Jakub Prokop, Martina Pecharová, Michael S. Engel & Romain Garrouste
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Meganisoptera was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itNeodonatoptera through timeFormations of the PermianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Martynov 1932