Pentazonia
Pentazonia is a taxonomic infraclass of millipedes containing the pill-millipedes (Oniscomorpha) which can roll into a ball and the order Glomeridesmida which cannot. Defining traits (apomorphies) include divided sternites, a labrum with single median tooth, and an enlarged pygidium on the hind-most body segment. Pentazonia is in the dominant millipede subclass Chilognatha which have a calcified exoskeleton and modified sperm-transferring legs in males. In contrast to the Helminthomorpha – the other Chilognathan infraclass, the sperm-transferring legs are located on posterior body segments and known as telopods. Pentazonians are relatively short-bodied, with between 13 and 21 body segments. The Pentazonia contains one extinct order, Amynilyspedida, often referred to the Oniscomorpha.

Ralf Janssen, Nikola-Michael Prpic and Wim GM Damen · CC BY 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ChilognathaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
Time the record covers
At least 406 million years on record.
The record covers at least 426.7 Mya to 20.45 Mya. No occurrence read is dated outside 432.9 Mya to 13.82 Mya.
26 occurrences of Pentazonia on record, most of them in the Carboniferous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
26 occurrences- Silurian4%
- Carboniferous62%
- Late Cretaceous19%
- Eocene12%
- Miocene4%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
5 formations
- Kladno Formation6
- Carbondale Formation · Kewanee Group5
- Joggins Formation · Cumberland Group1
- Fish Bed Formation · Glenbuck Group1
- Brooch Formation1
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The named rock units Pentazonia has been recovered from, largest first. Drawn where the rock is now, not where it formed. 12 occurrences have no formation on record, which is common for older literature.
Pentazonia and its relatives
CarboniferousWhere Pentazonia and its relatives were recorded in the Carboniferous.
Found in the same rock
1,405 taxa- Glomerida3 sites
- Palaeocampa anthrax3 sites
- Pleurojulus biornatus3 sites
- Ablator triguttatus2 sites
- Amynilyspes2 sites
- Amynilyspes fatimae2 sites
- Amynilyspes wortheni2 sites
- Andrognathus burmiticus2 sites
- Apidae2 sites
- Araneae2 sites
- Archaea paradoxa2 sites
- Blanziulus parriati2 sites
- Bradoponera meieri2 sites
- Cambalidea2 sites
- Carebara antiqua2 sites
- Clostes priscus2 sites
- Clya obscura2 sites
- Gesomyrmex hoernesi2 sites
- Glomeridesmidae2 sites
- Gnamptogenys europaea2 sites
- Hahniidae2 sites
- Heterochordeumatidae2 sites
- Hyleoglomeris groehni2 sites
- Lasius schiefferdeckeri2 sites
The Paleobiology Database records 17 sites for Pentazonia, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
11 references- Fritsch, 18996
- Checklist of the Myriapoda in Cretaceous Burmese amber and a correction of the Myriapoda identified by Zhang (2017)5
- A new species of the diplopod Amynilyspes (Oniscomorpha) from the Stephanian lagerstätte of Montceau-les-Mines, France3
- Systematics and functional morphology of oniscomorph millipedes (Arthropoda: Diplopoda) from the Carboniferous of North America3
- The oldest pill millipede fossil: A species of the Asiatic pill millipede genus Hyleoglomeris in Baltic amber (Diplopoda: Glomerida: Glomeridae)2
- Zosterogrammida, a new order of millipedes from the middle Silurian of Scotland and the upper Carboniferous of Euramerica2
- Millipeds from Dominican amber, with the description of two new species (Diplopoda: Siphonophoridae) of Siphonophora1
- The arthropod fauna of the Upper Carboniferous rocks of the Maritime Provinces1
- Canadian fossil insects, myriapods and arachnids, 3. Notes upon myriapods and arachnids found in sigillarian stumps in the Nova Scotia coal field1
- Archipolypoda, a subordinal type of spined myriapods from the Carboniferous formation1
- Footnotes and taxonomic names, in Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt1
Each of the 26 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Pentazonia
- Source
- Wikipedia/Commons
- Credit
- Ralf Janssen, Nikola-Michael Prpic and Wim GM Damen
- Licence
- CC BY 2.0
- Commercial use
- the licence allows it
No 3D scan of Pentazonia was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Pentazonia”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itChilognatha through timeFormations of the CarboniferousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Brandt 1833