Pilularia
Pilularia or pillworts is a genus of unusual ferns of family Marsileaceae distributed in North Temperate regions, Ethiopian mountains, and the southern hemisphere in Australia, New Zealand, and western South America.

Uncredited · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PlantaePBDB
Time the record covers
At least 56 million years on record.
The record covers at least 93.9 Mya to 37.71 Mya. No occurrence read is dated outside 100.5 Mya to 23.04 Mya.
2 occurrences of Pilularia on record, most of them in the Late Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
2 occurrences- Late Cretaceous50%
- Eocene13%
- Oligocene37%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
2 formations
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The named rock units Pilularia has been recovered from, largest first. Drawn where the rock is now, not where it formed.
Pilularia and its relatives
Late CretaceousWhere Pilularia and its relatives were recorded in the Late Cretaceous.
Found in the same rock
11 taxa- Arcellites disciformis1 site
- Balmeisporites1 site
- Chrysotheca1 site
- Chrysotheca diskoensis1 site
- Coptothyris1 site
- Microcarpolithes1 site
- Minerisporites1 site
- Molaspora lobata1 site
- Perotrilites1 site
- Regnellidium1 site
- Spermatites1 site
The Paleobiology Database records 1 site for Pilularia, and this reads all of them. The number is how many of those sites it was catalogued from too.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
2 references- Estudio palinológico de la Formación Loreto (Terciario Medio), Provincia de Magallanes, Chile1
- Megaspores and other fossils in the Dakota Formation (Cenomanian) of Iowa, (U.S.A.)1
Each of the 2 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Each entry links to the Paleobiology Database's own record for the work.
Media and terms
Media content1
Picture of Pilularia
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Pilularia was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Pilularia”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPlantae through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1753