Protohippus
Protohippus is an extinct three-toed genus of horse. It was roughly the size of a modern donkey. Fossil evidence suggests that it lived during the Late Miocene, from about 13.6 Ma to 5.3 Ma.

Jonathan Chen · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- EquiniPBDB
- Diet
- grazer, browserPBDB, for Equinae
- Life habit
- ground dwellingPBDB, for Equinae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 5.2 million years on record.
The record covers at least 12.5 Mya to 7.246 Mya. No occurrence read is dated outside 23.04 Mya to 4.7 Mya.
95 occurrences of Protohippus on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
95 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
15 formations
- Valentine Formation · Ogallala Group17
- Ash Hollow Formation · Ogallala Group15
- Fleming Formation8
- Ogallala Formation3
- Alachua Formation3
- Goliad Formation3
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The named rock units Protohippus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 33 occurrences have no formation on record, which is common for older literature.
Protohippus and its relatives
MioceneWhere Protohippus and its relatives were recorded in the Miocene.
Found in the same rock
727 taxa- Hipparionini23 sites
- Aelurodon ferox20 sites
- Epicyon saevus20 sites
- Camelidae19 sites
- Hippotherium19 sites
- Teleoceras major19 sites
- Aphelops megalodus18 sites
- Leptocyon vafer18 sites
- Calippus placidus16 sites
- Epicyon haydeni16 sites
- Teleoceras16 sites
- Aepycamelus15 sites
- Archaeolaginae15 sites
- Paratomarctus euthos15 sites
- Aelurodon taxoides14 sites
- Mylagaulidae14 sites
- Calippus proplacidus13 sites
- Calippus regulus13 sites
- Eucastor tortus13 sites
- Longirostromeryx wellsi13 sites
- Ustatochoerus medius13 sites
- Calippus martini12 sites
- Gomphotherium12 sites
- Paratomarctus temerarius12 sites
The Paleobiology Database records 95 sites for Protohippus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
32 references- Phylogenetic Interrelationships and Evolution of North American Late Equidae16
- Vertebrate biostratigraphy of the Ogallala Group in Nebraska12
- Calippus and Protohippus (Mammalia, Perissodactyla, Equidae) from the Miocene (Barstovian-early Hemphillian) of the Gulf Coastal Plain11
- Synonymies and reidentifications of North American fossil mammals7
- The Fossil Horses of the Texas Gulf Coastal Plain: A Revision7
- Cormohipparion and Hipparion (Mammalia, Perissodactyla, Equidae) from the Late Neogene of Florida6
- A Preliminary Report on the Miocene Vertebrate Faunas of Southeast Texas5
- The geology and paleontology of Ashfall Fossil Beds, a late Miocene (Clarendonian) mass-death assemblage, Antelope County and adjacent Knox County, Nebraska, USA2
- Road Log Neogene geology of the Española Basin, New Mexico2
- The Hottell Ranch rhino quarries (basal Ogallala: medial Barstovian), Banner County, Nebraska. Part I: Geologic setting, faunal lists, lower vertebrates2
- A faunal assemblage from the Lower Ash Hollow Formation (Neogene) of southern Nebraska2
- Miocene Equidae of the Texas Gulf Coastal Plain2
Each of the 95 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 20 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Protohippus
- Source
- Wikipedia/Commons
- Credit
- Jonathan Chen
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Protohippus was found in Smithsonian Open Access and Sketchfab.
MorphoSource was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Protohippus”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itEquini through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Leidy 1858)