Protosqualus
Protosqualus is an extinct genus of dogfish shark known from fossils found in Cretaceous rocks in Europe, Asia, Antarctica, Australia, and South America. The type species is Protosqualus sigei, which was found around an Albian-aged deposit in France. There are six species, which can be differentiated by distinct features in their teeth. Some species, such as P. barringtonensis, exhibit high levels of heterodonty. The oldest specimens are from the Speeton Clay Formation. Protosqualus teeth are quite common in the Grey Chalk deposit of England. The genus went extinct at the end of the Cretaceous during the Cretaceous–Paleogene extinction event, with the last species in the genus being Protosqualus argentinensis from southern Argentina as well as possibly being from earlier deposits in India.

Bubblesorg · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- SqualidaePBDB
- Diet
- carnivorePBDB, for Squaliformes
- Life habit
- nektonicPBDB, for Squaliformes
- Motility
- actively mobilePBDB, for Squaliformes
Time the record covers
At least 37 million years on record.
The record covers at least 121.4 Mya to 83.6 Mya. No occurrence read is dated outside 125.77 Mya to 72.2 Mya.
69 occurrences of Protosqualus on record, most of them in the Early Cretaceous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
69 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
11 formations
- Hunstanton Formation33
- Speeton Clay Formation7
- Labguva Formation4
- Jiesia Formation3
- Ferriby Formation2
- Mata Amarilla Formation2
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The named rock units Protosqualus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 13 occurrences have no formation on record, which is common for older literature.
Protosqualus and its relatives
Early CretaceousWhere Protosqualus and its relatives were recorded in the Early Cretaceous.
Found in the same rock
140 taxa- Squatina (Cretascyllium) cranei41 sites
- Gladioserratus aptiensis36 sites
- Synechodus dubrisiensis35 sites
- Cretascyliorhinus destombesi21 sites
- Heterodontus canaliculatus17 sites
- Archaeolamna kopingensis11 sites
- Eostriatolamia striatula11 sites
- Paraorthacodus recurvus11 sites
- Synechodus nitidus11 sites
- Squatirhina thiesi10 sites
- Leptostyrax macrorhiza9 sites
- Squalicorax falcatus9 sites
- Paranomotodon angustidens8 sites
- Squalicorax primaevus8 sites
- Anomotodon principalis7 sites
- Eostriatolamia subulata7 sites
- Heterodontus upnikensis7 sites
- Pseudoscapanorhynchus compressidens7 sites
- Dwardius woodwardi6 sites
- Orectoloboides parvulus6 sites
- Pteroscyllium ornatum6 sites
- Scapanorhynchus raphiodon6 sites
- Chiloscyllium greeni5 sites
- Cretorectolobus gracilis5 sites
The Paleobiology Database records 66 sites for Protosqualus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
14 references- Albian and Cenomanian selachian assemblages from north-east England40
- Re-evaluation of squaloid shark records from the Albian and Cenomanian of Lithuania8
- Les Elasmobranches de l'Albien inferieur et moyen (Cretace inferieur) de la Marne et de la Haute-Marne (France)5
- Sharks (Elasmobranchii: Euselachii) from the Late Cretaceous of France and the UK4
- Novel perspectives on Late Cretaceous elasmobranch diversity in the Southern Hemisphere high latitudes2
- Barremian and Aptian (Cretaceous) sharks and rays from Speeton, Yorkshire, NE England2
- Santonian-Campanian neoselachian faunas of the Upper Cretaceous Yezo Group in Nakagawa Town, Hokkaido, Japan1
- Elasmobranch (Chondrichthyes) assemblages from the Albian (Lower Cretaceous) of Ukraine1
- Marine vertebrates from the Santonian coastal carbonates of northwestern Germany - a tool for the reconstruction of a Proto-North Sea Basin intertidal dinosaur-exchange bridge1
- Marine vertebrates from the ‘Middle’ Cretaceous (Early Cenomanian) of South India1
- Additions to the selachian fauna of the Russian Cretaceous. 1. A new species of Protosqualus Cappetta, 1977 (Chondrichthyes, Squalidae)1
- Selachier (Pisces, Chondrichthyes) aus dem Cenomanium von Ascheloh am Teutoburger Wald (Nordrhein-Westfalen, NW-Deutschland)1
Each of the 69 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 2 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Protosqualus
- Source
- Wikipedia/Commons
- Credit
- Bubblesorg
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Protosqualus was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itSqualidae through timeFormations of the Early CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Cappetta 1977