Pseudopus
Pseudopus is a genus of anguid lizards that are native to Eurasia. One extant species remains, the sheltopusik, with four fossil species. They are the most robust members of subfamily Anguinae, with the largest species †P. pannonicus growing up to 2 metres in length. The oldest fossils of the group date to the Early Miocene, but there are possible Oligocene records.

(c) Ryan van Huyssteen, some rights reserved (CC BY-SA) · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- legless lizardPBDB
- Sits inside
- AnguinaePBDB
- Diet
- insectivore, carnivorePBDB, for Anguidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 22 million years on record.
The record covers at least 23.04 Mya to 129 ka. No occurrence read is dated outside 27.3 Mya to 11.7 ka.
111 occurrences of Pseudopus on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
111 occurrences- Oligocene3%
- Miocene62%
- Pliocene24%
- Pleistocene11%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
13 formations
- Upper Freshwater Molasse Formation · Molasse Group7
- Reuver clay Formation4
- Wiesbaden Formation2
- Most Formation2
- Ville Formation2
- Gaverdovsky Formation2
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The named rock units Pseudopus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 85 occurrences have no formation on record, which is common for older literature.
Pseudopus and its relatives
MioceneWhere Pseudopus and its relatives were recorded in the Miocene.
Found in the same rock
1,065 taxa- Lacerta37 sites
- Lacertidae28 sites
- Rana22 sites
- Colubridae21 sites
- Ophisaurus21 sites
- Colubrinae20 sites
- Vipera19 sites
- Natricinae18 sites
- Natrix17 sites
- Pelophylax16 sites
- Pelobates15 sites
- Elaphe14 sites
- Latonia gigantea14 sites
- Scincidae13 sites
- Natrix longivertebrata12 sites
- Anguis fragilis11 sites
- Serpentes11 sites
- Testudo11 sites
- Viperidae11 sites
- Coluber10 sites
- Eryx10 sites
- Ophisaurus spinari10 sites
- Ranidae10 sites
- Apodemus9 sites
The Paleobiology Database records 90 sites for Pseudopus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
45 references- Late Cainozoic localities of snakes and lizards of Ukraine14
- Revision of the anguine lizard Pseudopus laurillardi (Squamata, Anguidae) from the Miocene of Europe, with comments on paleoecology11
- The easternmost record of the largest anguine lizard that has ever lived – Pseudopus pannonicus (Squamata, Anguidae): new fossils from the late Neogene of Eastern Europe7
- Fossil squamate faunas from the Neogene of Hambach (northwestern Germany)6
- Late Turolian amphibians and reptiles from Brisighella (Northern Italy): preliminary report5
- Notes on the amphibian and reptilian fauna of the Polish Pliocene and Early Pleistocene5
- Amphibians and squamate reptiles from the late Miocene of Fălciu (Eastern Romania)4
- Amphibians and Squamates from the Miocene of Felsőtárkány Basin, N-Hungary4
- New material of Ophisaurus, Anguis and Pseudopus (Squamata, Anguidae, Anguinae) from the Miocene of the Czech Republic and Germany and systematic revision and palaeobiogeography of the Cenozoic Anguinae3
- The Early Miocene herpetofauna (Amphibia and Squamata) from Sibnica, Serbia3
- A new species of Pseudopus (Squamata, Anguidae) from the early Miocene of Northwest Bohemia (Czech Republic)3
- Lizards from the Late Miocene of Polgárdi (W-Hungary)3
Each of the 111 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 33 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Pseudopus
- Source
- Wikipedia/Commons
- Credit
- (c) Ryan van Huyssteen, some rights reserved (CC BY-SA)
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Pseudopus was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Pseudopus”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAnguinae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Merrem 1820