Ptychopteridae
The Ptychopteridae, phantom crane flies, are a small family of nematocerous Diptera. Superficially similar in appearance to other "tipuloid" families, they lack the ocelli of the Trichoceridae, the five-branched radial vein of the Tanyderidae, and the two anal veins that reach the wing margins of the Tipulidae. They are usually allied with the Tanyderidae based on similarities of the mesonotal suture, this group being called the Ptychopteromorpha.

Backslash · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- phantom crane flyPBDB
- Sits inside
- PtychopteromorphaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 196 million years on record.
The record covers at least 205.7 Mya to 8.725 Mya. No occurrence read is dated outside 211.18 Mya to 5.09 Mya.
138 occurrences of Ptychopteridae on record, most of them in the Late Jurassic. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
138 occurrences- Late Triassic1%
- Early Jurassic19%
- Middle Jurassic13%
- Late Jurassic34%
- Early Cretaceous22%
- Late Cretaceous2%
- Eocene7%
- Miocene1%
- Plioceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
32 formations
- Karabastau Formation26
- Itat Formation11
- Daohugou Formation10
- Zaza Formation6
- Posidonia Shale Formation5
- Sharteg Formation5
Loading coastlines…
The named rock units Ptychopteridae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 8 further formations hold fewer occurrences each and are not listed. 28 occurrences have no formation on record, which is common for older literature.
Ptychopteridae and its relatives
Late JurassicWhere Ptychopteridae and its relatives were recorded in the Late Jurassic.
Found in the same rock
3,799 taxa- Procramptonomyia kovalevi8 sites
- Ampliceps furcitibia7 sites
- Archirhyphus asiaticus7 sites
- Limoniidae7 sites
- Orthophlebiidae7 sites
- Oxycorynoides (Oxycorynoides) similis7 sites
- Panorpidium geinitzi7 sites
- Protoscelis jurassica7 sites
- Protoscelis longicornis7 sites
- Ampliceps dubius6 sites
- Anaxyelidae6 sites
- Codemus synaptoides6 sites
- Distenorrhinus (Distenorrhinus) pallidirostris6 sites
- Eoptychoptera simplex6 sites
- Liassothrips crassipes6 sites
- Panorpidium media6 sites
- Panorpidium minima6 sites
- Paramesopsocus adibi6 sites
- Polyanka minuta6 sites
- Praemacrochile ansorgei6 sites
- Protoscelis nitidicornis6 sites
- Rhagionidae6 sites
- Ampliceps dentitibia5 sites
- Ampliceps tenuirostris5 sites
The Paleobiology Database records 71 sites for Ptychopteridae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
45 references- New Eoptychopteridae (Diptera: Psychodomorpha) from the Jurassic of Asia27
- Revision of Eoptychopterinae (Diptera: Eoptychopteridae)9
- New Mesozoic psychodomorph dipteran insects from Siberia (Diptera: Eoptychopteridae, Ptychopteridae)9
- New true flies (Insecta: Diptera) from the Lower Cretaceous of southern England8
- Infraorders Psychodomorpha, Tipulomorpha and Culicomorpha8
- Five new species of Eoptychopteridae (Diptera) from the Mesozoic of Asia7
- A revision of the genus Eoptychopterina (Diptera: Eoptychopteridae)6
- Die Insektenfauna des Ostniedersachsischen Oberen Lias5
- New fossils of Eoptychopteridae (Diptera) from the Middle Jurassic of northeastern China3
- Eoptychopteridae (Diptera) from the Middle Jurassic of China3
- Bittacomorphinae (Diptera: Ptychopteridae) aus der alttertiären Fur-Formation (Moler) von Dänemark3
- Insect-bearing horizons in the type Purbeck and new Purbeck/Wealden flies (Diptera)3
Each of the 138 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 33 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ptychopteridae
- Source
- Wikipedia/Commons
- Credit
- Backslash
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Ptychopteridae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPtychopteromorpha through timeFormations of the Late JurassicThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Osten-Sacken 1862