Hymenoptera
Searched as Aculeata. The Paleobiology Database treats Hymenoptera as the accepted name.
Aculeata is an infraorder of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot sting, either retaining the ovipositor, or having lost it altogether. A large part of the clade is parasitic.

Uncredited · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- HymenopteridaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 469 million years on record.
The record covers at least 469.1 Mya to 11.7 ka. No occurrence read is dated outside 471.3 Mya to 0 Mya.
6,576 occurrences of Hymenoptera on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
6,576 occurrences- Ordovicianunder 1%
- Middle Triassicunder 1%
- Late Triassicunder 1%
- Early Jurassicunder 1%
- Middle Jurassic1%
- Late Jurassic6%
- Early Cretaceous11%
- Late Cretaceous13%
- Paleoceneunder 1%
- Eocene41%
- Oligocene6%
- Miocene16%
- Plioceneunder 1%
- Pleistocene1%
- Holocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
173 formations
- Florissant Formation440
- Karabastau Formation239
- Daohugou Formation172
- Yixian Formation · Jehol Group159
- Shanwang Formation146
- Bouldnor Formation · Solent Group136
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The named rock units Hymenoptera has been recovered from, largest first. Drawn where the rock is now, not where it formed. 149 further formations hold fewer occurrences each and are not listed. 3,477 occurrences have no formation on record, which is common for older literature.
Hymenoptera and its relatives
EoceneWhere Hymenoptera and its relatives were recorded in the Eocene.
Found in the same rock
8,102 taxa- Coleoptera16 sites
- Diptera12 sites
- Formicidae12 sites
- Culicoides filipalpis11 sites
- Curculionidae11 sites
- Serphites paradoxus10 sites
- Atriculicoides globosus8 sites
- Carabidae8 sites
- Ellinaphidinae8 sites
- Plecia foersteri8 sites
- Protoscelis jurassica8 sites
- Archirhyphus asiaticus7 sites
- Bibio7 sites
- Bibionidae7 sites
- Cydnus brevicollis7 sites
- Liassothrips crassipes7 sites
- Paramesopsocus adibi7 sites
- Peronehelea chrimikalydia7 sites
- Polyphaga7 sites
- Protoscelis longicornis7 sites
- Tipula7 sites
- Acrididae6 sites
- Aphanus pulchellus6 sites
- Atriculicoides6 sites
The Paleobiology Database records at least 335 sites for Hymenoptera, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
1,416 references- Die Ameisen des baltischen Bernsteins169
- Hymenoptera Apocrita of Mesozoic148
- The Fossil Ants of North America146
- The Parasitic Hymenoptera of the Baltic Amber142
- The parasitic Hymenoptera of the Tertiary of Florissant, Colorado123
- The Ants of the Baltic Amber115
- Les insectes fossiles des terrains oligocènes de France112
- Zhang, 198997
- Heer, 184994
- Modernisation of the Hymenoptera: ants, bees, wasps, and sawflies of the early Eocene Okanagan Highlands of western North America86
- A monograph of the Baltic amber bees and evolution of the Apoidea (Hymenoptera)84
- The wasps, bees and ants (Insecta: Vespida=Hymenoptera) from the Insect Limestone (Late Eocene) of the Isle of Wight, UK78
Each of the 6,576 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 1,404 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3
Picture of Hymenoptera
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Bulimina aculeata · l-cb
3D model
- Source
- Sketchfab
- Credit
- Shellie Luallin
- Licence
- CC Attribution
- Commercial use
- cannot be read from the licence given — check the record
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Aculeata”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itHymenopterida through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Scopoli 1763