Adephaga
The Adephaga are a suborder of beetles, and with more than 40,000 recorded species in 10 families, the second-largest of the four beetle suborders. Members of this suborder are collectively known as adephagans. The largest family is Carabidae which comprises most of the suborder with over 40,000 species. Adephaga also includes a variety of aquatic beetles, such as predaceous diving beetles and whirligig beetles.

Udo Schmidt from Deutschland · CC BY-SA 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- MetacoleopteraPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 251 million years on record.
The record covers at least 251.902 Mya to 11.7 ka. No occurrence read is dated outside 254.14 Mya to 0 Mya.
2,463 occurrences of Adephaga on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
2,463 occurrences- Permianunder 1%
- Early Triassicunder 1%
- Middle Triassicunder 1%
- Late Triassic2%
- Early Jurassic4%
- Middle Jurassicunder 1%
- Late Jurassic7%
- Early Cretaceous9%
- Late Cretaceous1%
- Paleoceneunder 1%
- Eocene9%
- Oligocene3%
- Miocene8%
- Pliocene2%
- Pleistocene52%
- Holocene1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
183 formations
- Kap København Formation236
- Highbury Silts and Sands Formation147
- Titusville Till Formation108
- Olympia beds Formation65
- Florissant Formation63
- Karabastau Formation56
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The named rock units Adephaga has been recovered from, largest first. Drawn where the rock is now, not where it formed. 159 further formations hold fewer occurrences each and are not listed. 740 occurrences have no formation on record, which is common for older literature.
Adephaga and its relatives
PleistoceneWhere Adephaga and its relatives were recorded in the Pleistocene.
Found in the same rock
7,260 taxa- Anostraca21 sites
- Schistomerus californense21 sites
- Dasyhelea antiqua20 sites
- Coleoptera19 sites
- Carabidae18 sites
- Curculionidae15 sites
- Hydrophilidae12 sites
- Thripidae12 sites
- Elateridae11 sites
- Olophrum rotundicolle11 sites
- Protoarrenurus convergens11 sites
- Memptus handlirschi10 sites
- Mesobaetis sibirica10 sites
- Orthoptera10 sites
- Stygeonectes jurassicus10 sites
- Formicidae9 sites
- Dyschirius8 sites
- Helophorus sempervarians8 sites
- Holocentropus affinis8 sites
- Omaliinae8 sites
- Panorpidium geinitzi8 sites
- Pycnoglypta8 sites
- Staphylinidae8 sites
- Stenus8 sites
The Paleobiology Database records at least 550 sites for Adephaga, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
367 references- Late Cenozoic insects of northern Eurasia288
- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland236
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK142
- Fossil Beetle Evidence for a Short Warm Interval near 40,000 yr B.P. at Titusville, Pennsylvania106
- Mezozoiskie zhestkokryiye98
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire67
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.65
- Beetles, Scarabaeida, in Pozdne-Mezozoyskie Nasekomye Vostochnogo Zabaykal'ya49
- Pleistoceńskie owady z Borysławia46
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA43
- Zhestkokrylye iz Yury Sibiri i zapadnoy Mongolii38
- Canadian fossil insects, 4. Additions to the coleopterous fauna of the interglacial clays of the Toronto district37
Each of the 2,463 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 355 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Adephaga
- Source
- Wikipedia/Commons
- Credit
- Udo Schmidt from Deutschland
- Licence
- CC BY-SA 2.0
- Commercial use
- the licence allows it
No 3D scan of Adephaga was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Adephaga”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMetacoleoptera through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Clairville 1806