Amphiesmenoptera
Amphiesmenoptera is an insect superorder, established by S. G. Kiriakoff, but often credited to Willi Hennig in his revision of insect taxonomy for two sister orders: Lepidoptera and Trichoptera (caddisflies). In 2017, a third fossil order was added to the group, the Tarachoptera.

Kropsoq · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PanorpidaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 316 million years on record.
The record covers at least 316.9 Mya to 11.7 ka. No occurrence read is dated outside 318.7 Mya to 0 Mya.
1,859 occurrences of Amphiesmenoptera on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
1,859 occurrences- Carboniferousunder 1%
- Permian5%
- Middle Triassicunder 1%
- Late Triassicunder 1%
- Early Jurassic3%
- Middle Jurassic1%
- Late Jurassic4%
- Early Cretaceous9%
- Late Cretaceous11%
- Paleoceneunder 1%
- Eocene46%
- Oligocene3%
- Miocene9%
- Plioceneunder 1%
- Pleistocene5%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
163 formations
- Florissant Formation59
- Amka Formation46
- Bouldnor Formation · Solent Group32
- Kap København Formation32
- Daohugou Formation27
- Koshelevka Formation24
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The named rock units Amphiesmenoptera has been recovered from, largest first. Drawn where the rock is now, not where it formed. 139 further formations hold fewer occurrences each and are not listed. 1,129 occurrences have no formation on record, which is common for older literature.
Amphiesmenoptera and its relatives
EoceneWhere Amphiesmenoptera and its relatives were recorded in the Eocene.
Found in the same rock
7,690 taxa- Coleoptera17 sites
- Plectrocnemia barbata13 sites
- Plectrocnemia lata13 sites
- Formicidae12 sites
- Lype sericea12 sites
- Yantaromyrmex geinitzi12 sites
- Ctenobethylus goepperti11 sites
- Formica flori11 sites
- Gesomyrmex hoernesi11 sites
- Holocentropus affinis11 sites
- Lepidoptera11 sites
- Trichoptera11 sites
- Curculionidae10 sites
- Diptera10 sites
- Holocentropus incertus10 sites
- Holocentropus vetustus10 sites
- Lasius schiefferdeckeri10 sites
- Prenolepis henschei10 sites
- Eocamponotus mengei9 sites
- Nylanderia pygmaea9 sites
- Plagiolepis klinsmanni9 sites
- Wormaldia aequalis9 sites
- Gryllidae8 sites
- Tetraponera simplex8 sites
The Paleobiology Database records 547 sites for Amphiesmenoptera, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
444 references- Die Trichopteren des baltischen Bernsteins357
- Wichard, 201363
- Melovye entomofauny basseyna r. Ul'i (zapadnoe Priokhot'e)46
- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland32
- Köcherfliegen des Baltischen Bernsteins 1. Marilia altrocki sp. n. (Trichoptera, Odontoceridae) der Bernsteinsammlung Bachofen-Echt32
- Die im Bernstein befindlichen Neuropteren der Vorwelt29
- The Khasurty fossil insect Lagerstätte28
- Paleontologiya cheshukerylykh i voprosy filogenii otrayada Papilionida24
- The Tertiary Insects of North America24
- Fossil caddisflies (Insecta: Trichoptera) from the Don Formation, Toronto, Ontario, and their use in paleoecology20
- Neue Untersuchungen über die fossilen Insekten mit Ergänzungen und Nachträgen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II Teil20
- Bernstein-Lepidopteren20
Each of the 1,859 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 432 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Amphiesmenoptera
- Source
- Wikipedia/Commons
- Credit
- Kropsoq
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Amphiesmenoptera was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPanorpida through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Kiriakoff 1948