Arctocyonidae
Arctocyonidae is an extinct polyphyletic family of placental mammals which lived from the Early Paleocene to the Early Eocene. They first appeared roughly 500,000 years after the Cretaceous-Paleogene extinction event, which wiped out the non-avian dinosaurs. They were initially regarded as creodonts, though have since been reassigned to an order of their own, the Arctocyonia. They were the most diverse during the Paleocene epoch, although a few such as Anacodon, Chriacus, and Thryptacodon, were present in the Early Eocene. All three genera would go extinct during the Early Eocene roughly 53.1 Ma. Some have suggested that arctocyonids are ancestral to modern-day artiodactyls, or that they form a sister group. However, more recent phylogenetic analyses suggest that arctocyonids may represent an artificial grouping of extinct ungulates, or that they might be an assemblage of unrelated placentals related to pangolins, pantodonts, and periptychids.

Dmitry Bogdanov · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- condylarthPBDB
- Sits inside
- ProcreodiPBDB
- Diet
- omnivorePBDB
- Life habit
- ground dwellingPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 17 million years on record.
The record covers at least 66 Mya to 48.07 Mya. No occurrence read is dated outside 72.2 Mya to 37.71 Mya.
920 occurrences of Arctocyonidae on record, most of them in the Paleocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
920 occurrences- Late Cretaceousunder 1%
- Paleocene80%
- Eocene19%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
52 formations
- Fort Union Formation187
- Willwood Formation139
- Hell Creek Formation132
- Nacimiento Formation79
- Ferris Formation36
- North Horn Formation29
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The named rock units Arctocyonidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 28 further formations hold fewer occurrences each and are not listed. 102 occurrences have no formation on record, which is common for older literature.
Arctocyonidae and its relatives
PaleoceneWhere Arctocyonidae and its relatives were recorded in the Paleocene.
Found in the same rock
910 taxa- Mesodma36 sites
- Stygimys kuszmauli34 sites
- Baioconodon nordicus33 sites
- Protungulatum donnae32 sites
- Tricentes subtrigonus30 sites
- Anisonchus sectorius29 sites
- Tetraclaenodon puercensis28 sites
- Mimatuta25 sites
- Procerberus formicarum25 sites
- Arctocyon ferox24 sites
- Oxyprimus erikseni24 sites
- Periptychus carinidens24 sites
- Chriacus baldwini23 sites
- Promioclaenus acolytus23 sites
- Ptilodus23 sites
- Stygimys gratus22 sites
- Ignacius frugivorus21 sites
- Neoplagiaulax21 sites
- Chriacus pelvidens20 sites
- Cimexomys minor20 sites
- Mimatuta morgoth20 sites
- Bessoecetor septentrionalis19 sites
- Protungulatum gorgun19 sites
- Thryptacodon australis19 sites
The Paleobiology Database records 505 sites for Arctocyonidae, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
124 references- Synonymies and reidentifications of North American fossil mammals104
- The Bug Creek problem and the Cretaceous-Tertiary transition at McGuire Creek, Montana85
- The Tiffanian Land-Mammal Age (middle and late Paleocene) in the northern Bighorn Basin, Wyoming74
- Stratigraphy and mammalian paleontology of the McCullough Peaks, northern Bighorn Basin, Wyoming: Implications for biochronology, basin development, and community reorganization across the Paleocene-Eocene boundary74
- Paleocene vertebrate paleontology of the San Juan Basin, New Mexico39
- A new important record of earliest Cenozoic mammalian history: geologic setting, Multituberculata, and Peradectia36
- The Clarkforkian Land-Mammal Age and Mammalian Faunal Composition Across the Paleocene-Eocene Boundary25
- Faunal Zonation of the Willwood Formation in the Central Bighorn Basin, Wyoming23
- The beginning of the Age of Mammals22
- Paleocene mammalian biostratigraphy of the Carbon Basin, southeastern Wyoming, and age constraints on local phases of tectonism15
- Paleocene (Puercan-Torrejonian) mammalian faunas of the North Horn Formation, central Utah15
- Earliest Eocene mammalian fauna from the Paleocene-Eocene Thermal Maximum at Sand Creek Divide, southern Bighorn Basin, Wyoming13
Each of the 920 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 112 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Arctocyonidae
- Source
- Wikipedia/Commons
- Credit
- Dmitry Bogdanov
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Arctocyonidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itProcreodi through timeFormations of the PaleoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Murray 1866