Belostomatidae
Belostomatidae is a family of freshwater hemipteran insects known as giant water bugs or colloquially as toe-biters, Indian toe-biters, electric-light bugs, alligator ticks, or alligator fleas. They are the largest insects in the order Hemiptera. There are about 170 species found in freshwater habitats worldwide, with more than 110 in the Neotropics, more than 20 in Africa, almost as many in the Nearctic, and far fewer elsewhere. These predators are typically encountered in freshwater ponds, marshes and slow-flowing streams. Most species are at least 2 cm (0.8 in) long, although smaller species, down to 0.9 cm (0.35 in), also exist. The largest are members of the genus Lethocerus, which can exceed 12 cm (4.5 in) and nearly reach the length of some of the largest beetles in the world. Giant water bugs are a popular food in parts of Asia.

Jhou5 · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- giant water bugPBDB
- Sits inside
- NepoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 205 million years on record.
The record covers at least 205.7 Mya to 11.7 ka. No occurrence read is dated outside 227.3 Mya to 0 Mya.
57 occurrences of Belostomatidae on record, most of them in the Late Jurassic. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
57 occurrences- Late Triassic2%
- Early Jurassic5%
- Middle Jurassic2%
- Late Jurassic37%
- Early Cretaceous25%
- Late Cretaceous2%
- Paleocene2%
- Eocene2%
- Oligocene5%
- Miocene16%
- Holocene4%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
18 formations
- Solnhofen Formation20
- Crato Formation · Santana Group7
- Shanwang Formation3
- Upper Freshwater-Molasse Formation2
- La Huérguina Formation2
- Sierra Madre Formation1
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The named rock units Belostomatidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 10 occurrences have no formation on record, which is common for older literature.
Belostomatidae and its relatives
Late JurassicWhere Belostomatidae and its relatives were recorded in the Late Jurassic.
Found in the same rock
1,548 taxa- Cymatophlebia longialata7 sites
- Isophlebia aspasia7 sites
- Scarabaeides deperditus7 sites
- Aeschnogomphus intermedius6 sites
- Chresmoda obscura6 sites
- Ditomoptera dubia6 sites
- Hexagenites cellulosus6 sites
- Lithoblatta lithophila6 sites
- Mesonepa primordialis6 sites
- Myrmicium schroeteri6 sites
- Pseudohydrophilus avitus6 sites
- Pycnophlebia speciosa6 sites
- Tarsophlebia eximia6 sites
- Actea sphinx5 sites
- Elcana lithophila5 sites
- Mesephemera procera5 sites
- Mesuropetala muensteri5 sites
- Stenophlebia latreillei5 sites
- Archipsyche eichstattensis4 sites
- Ephemeroptera4 sites
- Insecta4 sites
- Locustopsis germari4 sites
- Nannogomphus bavaricus4 sites
- Odonata4 sites
The Paleobiology Database records 45 sites for Belostomatidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
36 references- Les Heteroptera aquatiques fossiles, état actuel des connaissances (Heteroptera: Nepomorpha et Gerromorpha)6
- Jurassic insects from Solenhofen in the Carnegie Museum and the Museum of Comparative Zoology6
- A giant water bug from the Lower Cretaceous Crato Formation of Brazil (Heteroptera: Belostomatidae: Lethocerinae)2
- New fossil insect records from the Purbeck of Dorset and the Wealden of the Weald2
- British Upper Triassic and Lower Jurassic Heteroptera and Coleorrhyncha (Insecta: Hemiptera)2
- Zhang et al., 19942
- Fossil insects from the Tithonian "Solnhofener Plattenkalke" in the Museum of Natural History, Vienna2
- On some fossil Cryptocerata (Heteroptera, Insecta) with descriptions of a new genus et species2
- Late Quaternary insects of Rancho La Brea and McKittrick, California2
- Historical development of the infraorder Nepomorpha (Heteroptera)2
- Die Insecten aus dem lithographischen Schiefer2
- Sur les Insectes fossiles du Calcaire Lithographique de la Bavière, qui se trouvent au Musée Teyler2
Each of the 57 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 24 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Belostomatidae
- Source
- Wikipedia/Commons
- Credit
- Jhou5
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Belostomatidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itNepoidea through timeFormations of the Late JurassicThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Leach 1815