Ctenocystoidea
Ctenocystoidea is an extinct clade of echinoderms, which lived during the Cambrian and Ordovician periods. Unlike other echinoderms, ctenocystoids had bilateral symmetry, or were only very slightly asymmetrical. They are believed to be one of the earliest-diverging branches of echinoderms, with their bilateral symmetry a trait shared with other deuterostomes. Ctenocystoids were once classified in the taxon Homalozoa, also known as Carpoidea, alongside cinctans, solutes, and stylophorans. Homalozoa is now recognized as a polyphyletic group of echinoderms without radial symmetry. Ctenocystoids were geographically widespread during the Middle Cambrian, with one species surviving into the Late Ordovician.

Samuel Zamora, Imran A. Rahman, Andrew B. Smith · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- EchinodermataPBDB
- Life habit
- epifaunalPBDB
- Environment
- marinePBDB
Time the record covers
At least 47 million years on record.
The record covers at least 505.3 Mya to 458.2 Mya. No occurrence read is dated outside 514.5 Mya to 452.8 Mya.
34 occurrences of Ctenocystoidea on record, most of them in the Cambrian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
34 occurrences- Cambrian97%
- Ordovician3%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
12 formations
- Jince Formation15
- Lead Bell Shale Formation3
- Langston Formation3
- Usarzow Sandstone Formation3
- Ferrals Formation2
- Jbel Wawrmast Formation · Feijas Group2
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The named rock units Ctenocystoidea has been recovered from, largest first. Drawn where the rock is now, not where it formed.
Ctenocystoidea and its relatives
CambrianWhere Ctenocystoidea and its relatives were recorded in the Cambrian.
Found in the same rock
165 taxa- Ceratocystis perneri6 sites
- Etoctenocystis5 sites
- Stromatocystites pentangularis5 sites
- Ceratocystis4 sites
- Ctenocystis utahensis4 sites
- Etoctenocystis bohemica4 sites
- Lichenoides priscus4 sites
- Stromatocystites4 sites
- Akadocrinus3 sites
- Akadocrinus jani3 sites
- Cigara3 sites
- Conocoryphe3 sites
- Conocoryphe sulzeri3 sites
- Ctenocystis3 sites
- Gogia3 sites
- Hyolithes3 sites
- Jugoszovia archaeocyathoides3 sites
- Lobocephalina emmrichi3 sites
- Paradoxides (Eccaparadoxides)3 sites
- Paradoxides (Eccaparadoxides) pusillus3 sites
- Paradoxides (Hydrocephalus)3 sites
- Peronopsis3 sites
- Ptychoparia striata3 sites
- Acadoparadoxides2 sites
The Paleobiology Database records 29 sites for Ctenocystoidea, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
20 references- Evolutionary implications of a new transitional blastozoan echinoderm from the middle Cambrian of the Czech Republic8
- A new Cambrian stem-group echinoderm reveals the evolution of the anteroposterior axis2
- Asturicystis havliceki sp. nov. (Echinodermata, Homostelea) from the Middle Cambrian of Bohemia (Barrandian area, Czech Republic)2
- Sistemática, Anatomía, Estructura y Función de Ctenocystoidea (Echinodermata Carpoidea del Paleozoico inferior)2
- Révision biostratigraphique du Cambrien moyen du versant méridional de la Montagne Noire (Languedoc, France)2
- Chlupáč et al., 19982
- Biostratigraphy of the Albertella and Glossopleura zones (lower Middle Cambrian) of northern Utah and southern Idaho2
- Morphology and evolution of blastozoan echinoderms2
- Redescription of the Cambrian edrioasteroid Sprinkleoglobus spencensis n. comb. (Wen et al., 2019) from the Spence Shale (Utah, USA)1
- The youngest ctenocystoids from the Upper Ordovician of the United Kingdom and the evolution of the bilateral body plan in echinoderms1
- The Cambrian edrioasteroid Stromatocystites (Echinodermata): Systematics, palaeogeography, and palaeoecology1
- The ontogeny of Cinctans (stem-group Echinodermata) as revealed by a new genus, Graciacystis, from the middle Cambrian of Spain1
Each of the 34 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 8 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ctenocystoidea
- Source
- Wikipedia/Commons
- Credit
- Samuel Zamora, Imran A. Rahman, Andrew B. Smith
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Ctenocystoidea was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itEchinodermata through timeFormations of the CambrianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Robison and Sprinkle 1969