Dinocephalia
Dinocephalia is a clade of generally large-bodied therapsids that flourished during Middle Permian between approximately 275 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included carnivorous (Anteosauria), as well as herbivorous and omnivorous (Tapinocephalia) forms, serving as both dominant megaherbivores and apex predators. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania.

DiBgd at English Wikipedia · CC BY 2.5 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- EutherapsidaPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 14 million years on record.
The record covers at least 274.4 Mya to 259.857 Mya. No occurrence read is dated outside 298.9 Mya to 251.902 Mya.
175 occurrences of Dinocephalia on record, most of them in the Permian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
175 occurrences- Permian100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
15 formations
- Abrahamskraal Formation · Beaufort Group48
- Amanakskaya Formation14
- Rio do Rasto Formation · Passa Dois Group7
- Bolshekinelskaya Formation4
- San Angelo Formation · Pease River Group2
- Qingtoushan Formation2
Loading coastlines…
The named rock units Dinocephalia has been recovered from, largest first. Drawn where the rock is now, not where it formed. 84 occurrences have no formation on record, which is common for older literature.
Dinocephalia and its relatives
PermianWhere Dinocephalia and its relatives were recorded in the Permian.
Found in the same rock
204 taxa- Anteosaurus magnificus10 sites
- Struthiocephalus whaitsi6 sites
- Titanosuchus ferox6 sites
- Anteosaurus5 sites
- Bashkyroleter mesensis5 sites
- Bradysaurus seeleyi5 sites
- Mesenosaurus romeri5 sites
- Syodon5 sites
- Tapinocephalidae5 sites
- Alrausuchus tagax4 sites
- Bradysaurus4 sites
- Jonkeria4 sites
- Nyctiphruretus acudens4 sites
- Syodon biarmicum4 sites
- Bradysaurus baini3 sites
- Deuterosaurus3 sites
- Dicynodon3 sites
- Discordichthys spinifer3 sites
- Emydops3 sites
- Enosuchus breviceps3 sites
- Glanosuchus macrops3 sites
- Platysomus biarmicus3 sites
- Tapinocephalia3 sites
- Actinopterygii2 sites
The Paleobiology Database records 72 sites for Dinocephalia, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
28 references- The fauna of the Tapinocephalus Zone (Beaufort Beds of the Karoo)64
- The distribution of the Karroo vertebrate fauna32
- Eotherapsids from the East European Placket (Late Permian)29
- Systematics of the Anteosauria (Therapsida: Dinocephalia)8
- A tapinocephalid dinocephalian (Synapsida, Therapsida) from the Rio do Rasto Formation (Parana Basin, Brazil): Taxonomic, ontogenetic and biostratigraphic considerations5
- Upper Permian vertebrates and their sedimentological context in the South Urals, Russia5
- A dinocephalian therapsid fauna on the Ecca–Beaufort contact in Eastern Cape Province, South Africa3
- Youngest dinocephalian fossils extend the Tapinocephalus Zone, Karoo Basin, South Africa2
- Type and reference sections of the Middle and Upper Permian of the Volga and Kama River Regions: A Field Guidebook of XVIII International Congress on Carboniferous and Permian2
- Tapinocephalids (Therapsida: Dinocephalia) from the Permian Madumabisa Mudstone Formation (Lower Karoo, Mid-Zambezi Basin) of southern Zambia2
- Cranial morphology and evolution of Permian Dinomorpha (Eotherapsida) of Eastern Europe2
- The most primitive lower tetrapod fauna in China2
Each of the 175 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 16 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Dinocephalia
- Source
- Wikipedia/Commons
- Credit
- DiBgd at English Wikipedia
- Licence
- CC BY 2.5
- Commercial use
- the licence allows it
No 3D scan of Dinocephalia was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itEutherapsida through timeFormations of the PermianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Seeley 1894)