Ichneumonidae
The Ichneumonidae, also known as ichneumon wasps, ichneumonid wasps, ichneumonids, or Darwin wasps, are a family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 25,000 species described as of 2016. However, this likely represents less than a quarter of their true richness as reliable estimates are lacking, along with much of the most basic knowledge about their ecology, distribution, and evolution. It is estimated that there are more species in this family than there are species of birds and mammals combined. Ichneumonid wasps, with very few exceptions, attack the immature stages of holometabolous insects and spiders, eventually killing their hosts. They play an important role as regulators of insect populations, both in natural and semi-natural systems, making them promising agents for biological control.

Uncredited · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- ichneumon waspPBDB
- Sits inside
- IchneumonoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 124 million years on record.
The record covers at least 125.77 Mya to 774 ka. No occurrence read is dated outside 132.6 Mya to 129 ka.
473 occurrences of Ichneumonidae on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
473 occurrences- Early Cretaceous11%
- Late Cretaceous8%
- Paleoceneunder 1%
- Eocene55%
- Oligocene10%
- Miocene9%
- Pliocene1%
- Pleistocene6%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
53 formations
- Florissant Formation85
- Fur Formation26
- Bouldnor Formation · Solent Group22
- Miyajima Formation · Shiobara Group17
- Ola Formation16
- Niveau du gypse d'Aix Formation16
Loading coastlines…
The named rock units Ichneumonidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 29 further formations hold fewer occurrences each and are not listed. 119 occurrences have no formation on record, which is common for older literature.
Ichneumonidae and its relatives
EoceneWhere Ichneumonidae and its relatives were recorded in the Eocene.
Found in the same rock
5,770 taxa- Formicidae20 sites
- Coleoptera11 sites
- Curculionidae10 sites
- Braconidae9 sites
- Mycetophilidae9 sites
- Prenolepis henschei9 sites
- Acrididae8 sites
- Anisoptera8 sites
- Gesomyrmex hoernesi8 sites
- Lepidoptera8 sites
- Nylanderia pygmaea8 sites
- Tetraponera simplex8 sites
- Yantaromyrmex geinitzi8 sites
- Aphididae7 sites
- Bibionidae7 sites
- Ctenobethylus goepperti7 sites
- Lasius schiefferdeckeri7 sites
- Plagiolepis kuenowi7 sites
- Aphanus pulchellus6 sites
- Bibio6 sites
- Ceratopogon6 sites
- Diapriinae6 sites
- Diptera6 sites
- Eohelea sinuosa6 sites
The Paleobiology Database records 172 sites for Ichneumonidae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
124 references- The parasitic Hymenoptera of the Tertiary of Florissant, Colorado71
- High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)23
- Les insectes fossiles des terrains oligocènes de France20
- Fossil Darwin wasps (Hymenoptera, Ichneumonidae) from the Middle Pleistocene Shiobara Group, Tochigi Prefecture, Japan17
- New ichneumonids (Insecta: Hymenoptera: Ichneumonidae) from the Lower Cretaceous of north China16
- A new subfamily of ichneumon wasps (Insecta: Hymenoptera: Ichneumonidae) from the Upper Cretaceous of the Russian far east16
- The wasps, bees and ants (Insecta: Vespida=Hymenoptera) from the Insect Limestone (Late Eocene) of the Isle of Wight, UK13
- A new subfamily of Ichneumonids from the Lower Cretaceous of Transbaikalia and Mongolia (Insecta: Hymenoptera: Ichneumonidae)12
- Ueber alte und neue fossile Hymenopterenfunde aus den Tertiaren Ablagerungen von Rott am Siebengebirge11
- Fossil Arthropods in the British Museum, V. Oligocene Hymenoptera from the Isle of Wight11
- New data on ichneumon wasps of the subfamily Pherhombinae (Hymenoptera, Ichneumonidae) in Baltic amber with descriptions of three new species9
- Fossil ichneumon wasps (Hymenoptera: Ichneumonidae) from Biamo (Russia), Oligocene9
Each of the 473 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 112 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ichneumonidae
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
No 3D scan of Ichneumonidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itIchneumonoidea through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1802