Ilybius
Ilybius is a large genus of predatory aquatic beetles in the family Dytiscidae. The genus is native to the Palearctic, the Near East, the Nearctic, and North Africa. 70 species has been described from this genus:Ilybius aenescens Thomson, 1870 Ilybius albarracinensis (Fery, 1986) Ilybius angustior (Gyllenhal, 1808) Ilybius anjarum Nilsson, 1999 Ilybius apicalis Sharp, 1873 Ilybius ater (De Geer, 1774) Ilybius austrodiscors (Larson, 1996) Ilybius balkei (Fery & Nilsson, 1993) Ilybius bedeli (Zaitzev, 1908) Ilybius biguttulus (Germar, 1824) Ilybius boryslavicus Lomnicki, 1894 Ilybius chalconatus (Panzer, 1796) Ilybius chishimanus Kôno, 1944 Ilybius churchillensis Wallis, 1939 Ilybius cinctus Sharp, 1878 Ilybius confertus (LeConte, 1861) Ilybius confusus Aubé, 1838 Ilybius crassus Thomson, 1856 Ilybius dettneri (Fery, 1986) Ilybius discedens Sharp, 1882 Ilybius discors (LeConte, 1861) Ilybius erichsoni (Gemminger & Harold, 1868) Ilybius euryomus (Larson, 1996) Ilybius fenestratus (Fabricius, 1781) Ilybius fraterculus LeConte, 1862 Ilybius fuliginosus (Fabricius, 1792) Ilybius gagates (Aubé, 1838) Ilybius guttiger (Gyllenhal, 1808) Ilybius hozgargantae (Burmeister, 1983) Ilybius hulae (Wewalka, 1984) Ilybius hypomelas (Mannerheim, 1843) Ilybius ignarus (LeConte, 1862) Ilybius incarinatus Zimmermann, 1928 Ilybius jaechi (Fery & Nilsson, 1993) Ilybius jimzim (Larson, 1996) Ilybius lagabrunensis (Schizzerotto & Fery, 1990) Ilybius larsoni (Fery & Nilsson, 1993) Ilybius lateralis (Gebler, 1832) Ilybius lenensis Nilsson, 2000 Ilybius lenkoranensis (Fery & Nilsson, 1993) Ilybius lineellus (LeConte, 1861) Ilybius meridionalis Aubé, 1837 Ilybius montanus (Stephens, 1828) Ilybius nakanei Nilsson, 1994 Ilybius neglectus (Erichson, 1837) Ilybius oblitus Sharp, 1882 Ilybius obtusus Sharp, 1882 Ilybius opacus (Aubé, 1837) Ilybius ovalis Gschwendtner, 1934 Ilybius pederzanii (Fery & Nilsson, 1993) Ilybius picipes (Kirby, 1837) Ilybius pleuriticus LeConte, 1850 Ilybius poppiusi Zaitzev, 1907 Ilybius pseudoneglectus (Franciscolo, 1972) Ilybius quadriguttatus (Lacordaire, 1835) Ilybius quadrimaculatus Aubé, 1838 Ilybius samokovi (Fery & Nilsson, 1993) Ilybius satunini (Zaitzev, 1913) Ilybius similis Thomson, 1856 Ilybius subaeneus Erichson, 1837 Ilybius subtilis (Erichson, 1837) Ilybius thynias Fery & Przewozny, 2011 Ilybius vancouverensis (Leech, 1937) Ilybius vandykei (Leech, 1942) Ilybius verisimilis (Brown, 1932) Ilybius vittiger (Gyllenhal, 1827) Ilybius walsinghami (Crotch, 1873) Ilybius wasastjernae (C.R.Sahlberg, 1824) Ilybius wewalkai (Fery & Nilsson, 1993)

James K. Lindsey · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- AgabinaePBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 33 million years on record.
The record covers at least 33.9 Mya to 129 ka. No occurrence read is dated outside 37.71 Mya to 11.7 ka.
28 occurrences of Ilybius on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
28 occurrences- Eocene7%
- Pliocene11%
- Pleistocene82%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
10 formations
- Kap København Formation6
- Tsubusagawa Formation3
- Highbury Silts and Sands Formation3
- Mizozono Formation2
- Olympia beds Formation2
- Bouldnor Formation · Solent Group2
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The named rock units Ilybius has been recovered from, largest first. Drawn where the rock is now, not where it formed. 6 occurrences have no formation on record, which is common for older literature.
Ilybius and its relatives
PleistoceneWhere Ilybius and its relatives were recorded in the Pleistocene.
Found in the same rock
1,247 taxa- Carabidae12 sites
- Bledius10 sites
- Curculionidae10 sites
- Gyrinus10 sites
- Stenus10 sites
- Aphodius9 sites
- Donacia9 sites
- Dytiscidae9 sites
- Staphylinidae9 sites
- Agabus8 sites
- Bembidion8 sites
- Chrysomelidae8 sites
- Amara7 sites
- Coleoptera7 sites
- Helophorus7 sites
- Plateumaris7 sites
- Aleocharinae6 sites
- Chrysolina6 sites
- Dyschirius6 sites
- Hydrobius fuscipes6 sites
- Hydroporus6 sites
- Lepidoptera6 sites
- Pterostichus6 sites
- Aegialia (Psammoporus) terminalis5 sites
The Paleobiology Database records 25 sites for Ilybius, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
14 references- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland6
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK3
- Fossil insects and their paleoenvironments of the Pliocene Tsubusagawa Formation in Ajimu Basin, Oita Prefecture, Kyushu, Japan3
- The beetle (Coleoptera) fauna of the Insect Limestone (late Eocene), Isle of Wight, southern England2
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.2
- Late Pleistocene insects from the Mizozono Formation in Yoshimatsu-cho, Kagoshima Prefecture, Japan2
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian2
- A Late Pleistocene fauna and flora from Upton Warren, Worcestershire2
- Pleistocene fossil insects from Tokachi, Hokkaido, north Japan1
- Middle Pleistocene fossil insects from the Saijo Formation of Higashi-hiroshima City, Hiroshima Prefecture, Japan1
- Middle Pleistocene fossil insects from the Hirabaru Formation of Kitsuki City, Oita Prefecture, Kyushu, Japan1
- Early Pleistocene ground beetles (Coleoptera: Carabidae) from the Ookui Formation in Nagano Prefecture, central Japan, and their biogeographical and paleoenvironmental significance1
Each of the 28 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 2 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ilybius
- Source
- Wikipedia/Commons
- Credit
- James K. Lindsey
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Ilybius”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAgabinae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Erichson 1832