Marrella
Marrella is an extinct genus of marrellomorph arthropod known from the Middle Cambrian of North America and Asia. It is the most common animal represented in the Burgess Shale of British Columbia, Canada, with tens of thousands of specimens collected. Much rarer remains are also known from deposits in China.

Jean-Bernard Caron · CC0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- MarrellidaPBDB
- Diet
- detritivorePBDB
- Life habit
- nektobenthicPBDB
- Motility
- actively mobilePBDB
Time the record covers
At most 18 million years on record.
Every occurrence read is dated to somewhere between 514.5 Mya and 497 Mya, and nothing in the dating separates them, so that window is what the record shows rather than a span.
31 occurrences of Marrella on record, most of them in the Cambrian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
31 occurrences- Cambrian100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
4 formations
- Burgess Shale Formation · Chancellor Group28
- Kaili Formation1
- Stephen Formation1
- Balang Formation1
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The named rock units Marrella has been recovered from, largest first. Drawn where the rock is now, not where it formed.
Marrella and its relatives
CambrianWhere Marrella and its relatives were recorded in the Cambrian.
Found in the same rock
275 taxa- Diraphora26 sites
- Hazelia26 sites
- Liangshanella26 sites
- Ehmaniella burgessensis25 sites
- Selkirkia columbia25 sites
- Wiwaxia corrugata25 sites
- Morania24 sites
- Pentagnostus praecurrens24 sites
- Sidneyia24 sites
- Yohoia24 sites
- Burgessia23 sites
- Canadaspis perfecta23 sites
- Ottoia prolifica23 sites
- Scenella amii23 sites
- Waptia fieldensis23 sites
- Haplophrentis carinatus22 sites
- Laggania22 sites
- Naraoia compacta22 sites
- Micromitra21 sites
- Pollingeria grandis21 sites
- Hurdia20 sites
- Trilobita20 sites
- Lingulella19 sites
- Molaria19 sites
The Paleobiology Database records 30 sites for Marrella, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
6 references- Paleoecology of the Greater Phyllopod Bed community, Burgess Shale25
- The Burgess Shale and associated Cambrian formations west of the Fossil Gully Fault Zone on Mount Stephen, British Columbia2
- A Cambrian soft-bodied biota after the first Phanerozoic mass extinction1
- Paleocommunity analysis of the Burgess Shale Tulip Beds, Mount Stephen, British Columbia: Comparison with the Walcott Quarry and implications for community variation in the Burgess Shale1
- A new phyllopod bed-like assemblage from the Burgess Shale of the Canadian Rockies1
- Basal Middle Cambrian short-stalked eorinoids from the Kali Biota: Guizhou Province, China1
Each of the 31 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Each entry links to doi.org. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Marrella
- Source
- Wikipedia/Commons
- Credit
- Jean-Bernard Caron
- Licence
- CC0
- Commercial use
- the licence allows it
No 3D scan of Marrella was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Marrella”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMarrellida through timeFormations of the CambrianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Walcott 1912