Milacidae
Milacidae is a family of air-breathing, keeled, land slugs. These are shell-less terrestrial gastropod mollusks in the superfamily Parmacelloidea.

Michal Maňas · CC BY 2.5 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ParmacelloideaPBDB
- Life habit
- epifaunalPBDB, for Gastropoda
- Motility
- actively mobilePBDB, for Gastropoda
- Vision
- limitedPBDB, for Gastropoda
- Environment
- terrestrialPBDB
Time the record covers
At least 36 million years on record.
The record covers at least 37.71 Mya to 774 ka. No occurrence read is dated outside 41.03 Mya to 11.7 ka.
24 occurrences of Milacidae on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
24 occurrences- Eocene4%
- Miocene63%
- Pliocene17%
- Pleistocene16%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
2 formations
Loading coastlines…
The named rock units Milacidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 13 occurrences have no formation on record, which is common for older literature.
Milacidae and its relatives
MioceneWhere Milacidae and its relatives were recorded in the Miocene.
Found in the same rock
448 taxa- Agriolimacidae12 sites
- Cochlostoma esuanum10 sites
- Limax fossilis10 sites
- Melanopsis narzolina10 sites
- Parmacella10 sites
- Saccoia globosa10 sites
- Saccoia oryza10 sites
- Truciella ballesioi10 sites
- Craspedopoma conoidale9 sites
- Lucilla miocaenica9 sites
- Melanoides curvicosta9 sites
- Nordsieckia pontica9 sites
- Theodoxus mutinensis9 sites
- Acicula lineata8 sites
- Chiroptera8 sites
- Clausilia (Clausilia) baudoni8 sites
- Helicodiscus roemeri8 sites
- Theodoxus doderleini8 sites
- Anguinae7 sites
- Apodemus atavus7 sites
- Apodemus gudrunae7 sites
- Centralomys benericettii7 sites
- Dreissena7 sites
- Glis minor7 sites
The Paleobiology Database records 24 sites for Milacidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
10 references- Late Messinian continental and Lago-Mare gastropods from the Tertiary Piedmont Basin, NW Italy8
- Moluscos continentales del Plio-Pleistoceno de la cuenca de Guadix-Baza7
- Late Messinian mollusks and vertebrates from Moncucco Torinese, north-western Italy. Paleoecological and paleoclimatological implications2
- Opole (Poland) - a key locality for middle Miocene terrestrial mollusc faunas1
- Early Miocene gastropod and ectothermic vertebrate remains from the Lesvos Petrified Forest (Greece)1
- Terrestrial, freshwater and brachyhaline Gastropoda from the Lower Miocene deposits of Oberdorf (Styria, Austria)1
- Synopsis of the Late Miocene mollusc fauna of the classical sections Richardhof and Eichkogel in the Vienna Basin (Austria, Pannonian, MN 9-MN11)1
- Tephrochronology and tephrostratigraphy of two Pleistocene continental fossiliferous successions from central Italy1
- The Creechbarrow Limestone - its biota and correlation1
- Dritter Beitrag zur Kenntnis des Miocäns von Oppeln i. Schl1
Each of the 24 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Milacidae
- Source
- Wikipedia/Commons
- Credit
- Michal Maňas
- Licence
- CC BY 2.5
- Commercial use
- the licence allows it
No 3D scan of Milacidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Milacidae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itParmacelloidea through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Ellis 1926