Nematocera
The Nematocera are a suborder of elongated flies with thin, segmented antennae and mostly aquatic larvae. This group is paraphyletic, containing all flies except for those of the suborder Brachycera, which includes species such as the housefly or the common fruit fly. Thus, the equivalent clade to Nematocera would be the whole of Diptera, with Brachycera as a "subclade". Families in Nematocera include mosquitoes, crane flies, gnats, black flies, and multiple families commonly known as midges.

Muhammad Mahdi Karim · GFDL 1.2 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- DipteraPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 245 million years on record.
The record covers at least 245.19 Mya to 11.7 ka. No occurrence read is dated outside 247 Mya to 0 Mya.
1,553 occurrences of Nematocera on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
1,553 occurrences- Middle Triassic1%
- Late Triassicunder 1%
- Early Jurassic8%
- Middle Jurassic3%
- Late Jurassic14%
- Early Cretaceous11%
- Late Cretaceous4%
- Paleoceneunder 1%
- Eocene44%
- Oligocene5%
- Miocene7%
- Plioceneunder 1%
- Pleistoceneunder 1%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
101 formations
- Florissant Formation93
- Bouldnor Formation · Solent Group74
- Karabastau Formation65
- Daohugou Formation59
- Glushkovo Formation50
- Posidonia Shale Formation37
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The named rock units Nematocera has been recovered from, largest first. Drawn where the rock is now, not where it formed. 77 further formations hold fewer occurrences each and are not listed. 700 occurrences have no formation on record, which is common for older literature.
Nematocera and its relatives
EoceneWhere Nematocera and its relatives were recorded in the Eocene.
Found in the same rock
7,545 taxa- Coleoptera14 sites
- Diptera14 sites
- Tipula11 sites
- Formicidae10 sites
- Panorpidium geinitzi9 sites
- Panorpidium minima9 sites
- Bibionidae8 sites
- Mycetophilidae8 sites
- Orthoptera8 sites
- Protoscelis jurassica8 sites
- Archirhyphus asiaticus7 sites
- Curculionidae7 sites
- Eocamponotus mengei7 sites
- Formica flori7 sites
- Gesomyrmex hoernesi7 sites
- Holocentropus affinis7 sites
- Insecta7 sites
- Lepidoptera7 sites
- Liassothrips crassipes7 sites
- Necrotaulius parvulus7 sites
- Plecia7 sites
- Protoscelis longicornis7 sites
- Aphaenogaster mayri6 sites
- Archaea paradoxa6 sites
The Paleobiology Database records 381 sites for Nematocera, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
364 references- Krzemińska et al., 2009109
- Crane-flies of the Baltic Amber (Diptera)93
- Tertiary Tipulidae, with special reference to those of Florissant, Colorado64
- Monographie des Tipulidae et des Dixidae de l'ambre de la Baltique46
- True flies (Insecta: Diptera) from the late Eocene insect limestone (Bembridge Marls) of the Isle of Wight, England, UK43
- Die Insektenfauna des Ostniedersachsischen Oberen Lias37
- New Eoptychopteridae (Diptera: Psychodomorpha) from the Jurassic of Asia27
- Infraorders Psychodomorpha, Tipulomorpha and Culicomorpha27
- Ormosia Rondani crane flies (Diptera, Limoniidae) from the Baltic amber24
- Neue Untersuchungen über die fossilen Insekten mit Ergänzungen und Nachträgen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II Teil24
- Limoniidae (Diptera) in the Upper Jurassic of Shar Teg, Mongolia21
- Rhabdomastix Crane Flies (Diptera: Limoniidae) in Baltic Amber (Eocene)18
Each of the 1,553 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 352 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Nematocera
- Source
- Wikipedia/Commons
- Credit
- Muhammad Mahdi Karim
- Licence
- GFDL 1.2
- Commercial use
- the licence allows it
No 3D scan of Nematocera was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Nematocera”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itDiptera through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1825