Opiliones
The Opiliones are an order of arachnids, colloquially known as harvestmen, harvesters, daddy long legs, or granddaddy long legs. Over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014.

Bruce Marlin · CC BY-SA 2.5 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- harvestmanPBDB
- Sits inside
- StomothecataPBDB
- Life habit
- solitaryPBDB, for Arachnida
- Motility
- actively mobilePBDB, for Arachnida
- Environment
- terrestrialPBDB, for Arachnida
Time the record covers
At least 390 million years on record.
The record covers at least 410.62 Mya to 20 Mya. No occurrence read is dated outside 413.02 Mya to 13.82 Mya.
123 occurrences of Opiliones on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
123 occurrences- Devonian2%
- Carboniferous12%
- Middle Jurassicunder 1%
- Late Jurassic2%
- Early Cretaceous2%
- Late Cretaceous28%
- Eocene47%
- Oligocene2%
- Miocene4%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
10 formations
- Carbondale Formation · Kewanee Group4
- Daohugou Formation3
- Florissant Formation3
- Niveau du gypse d'Aix Formation2
- Commentry Shales Formation2
- Messel Formation2
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The named rock units Opiliones has been recovered from, largest first. Drawn where the rock is now, not where it formed. 103 occurrences have no formation on record, which is common for older literature.
Opiliones and its relatives
EoceneWhere Opiliones and its relatives were recorded in the Eocene.
Found in the same rock
3,930 taxa- Araneae7 sites
- Leiobunum longipes7 sites
- Ctenobethylus goepperti6 sites
- Dicranopalpus ramiger6 sites
- Caddo dentipalpus5 sites
- Dasyhelea5 sites
- Gesomyrmex hoernesi5 sites
- Lasaeola infulata5 sites
- Lasius schiefferdeckeri5 sites
- Monogedania clunipes5 sites
- Nylanderia pygmaea5 sites
- Pachycondyla succinea5 sites
- Plagiolepis klinsmanni5 sites
- Prenolepis henschei5 sites
- Tetraponera simplex5 sites
- Trichoneura (Trichoneura) vulgaris5 sites
- Yantaromyrmex geinitzi5 sites
- Archaea paradoxa4 sites
- Architarbus rotundatus4 sites
- Brachycera4 sites
- Brachypogon (Isohelea) prominulus4 sites
- Bradoponera meieri4 sites
- Ceratopogon4 sites
- Clostes priscus4 sites
The Paleobiology Database records 53 sites for Opiliones, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
51 references- An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber10
- Baltic amber harvestmen (Opiliones) from Polish collections10
- Footnotes and taxonomic names, in Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt10
- First eupnoid harvestmen (Arachnida: Opiliones: Eupnoi) from mid-Cretaceous Kachin amber, with notes on sexual dimorphism in Halitherses grimaldii (Arachnida: Opiliones: Dyspnoi)7
- Laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber7
- Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt7
- Fossil harvestmen (Arachnida, Opiliones) from Bitterfeld amber6
- Opilioniden im Bernstein5
- Four new laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber4
- Iridescent harvestmen (Arachnida: Opiliones: Sclerosomatidae) from the Eocene of Messel, Germany2
- A new species of Laniatores (Arachnida: Opiliones) from Eocene Baltic amber with notes on the evolution of Insidiatores2
- Two laniatorid harvestmen (Opiliones: Cladonychiidae) from Eocene Baltic amber2
Each of the 123 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 39 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Opiliones
- Source
- Wikipedia/Commons
- Credit
- Bruce Marlin
- Licence
- CC BY-SA 2.5
- Commercial use
- the licence allows it
No 3D scan of Opiliones was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Opiliones”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itStomothecata through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Sundevall 1833