Priapulida
Priapulida, sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility, because their general shape and their extensible spiny introvert (eversible) proboscis may resemble the shape of a human penis.

Shunkina Ksenia · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- priapulid wormPBDB
- Sits inside
- ScalidophoraPBDB
- Diet
- carnivorePBDB
- Life habit
- shallow infaunalPBDB
- Motility
- facultatively mobilePBDB
- Vision
- blindPBDB
Time the record covers
At least 215 million years on record.
The record covers at least 525.5 Mya to 309.8 Mya. No occurrence read is dated outside 538.8 Mya to 307.4 Mya.
386 occurrences of Priapulida on record, most of them in the Cambrian. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
386 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
64 formations
- Burgess Shale Formation · Chancellor Group87
- Yuanshan Formation77
- Yu'anshan Formation58
- Qiongzhusi Formation18
- Balang Formation13
- Ella Island Formation12
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The named rock units Priapulida has been recovered from, largest first. Drawn where the rock is now, not where it formed. 40 further formations hold fewer occurrences each and are not listed. 2 occurrences have no formation on record, which is common for older literature.
Priapulida and its relatives
CambrianWhere Priapulida and its relatives were recorded in the Cambrian.
Found in the same rock
1,517 taxa- Hazelia39 sites
- Mafangscolex sinensis32 sites
- Arthropoda29 sites
- Selkirkia29 sites
- Trilobita29 sites
- Heliomedusa orienta28 sites
- Liangshanella28 sites
- Lingulella28 sites
- Wiwaxia corrugata28 sites
- Diraphora27 sites
- Selkirkia columbia27 sites
- Ehmaniella burgessensis26 sites
- Isoxys auritus26 sites
- Kunmingella douvillei26 sites
- Ottoia prolifica26 sites
- Burgessia25 sites
- Haplophrentis carinatus25 sites
- Marrella splendens25 sites
- Micromitra25 sites
- Misszhouia longicaudata25 sites
- Morania25 sites
- Naraoia compacta25 sites
- Pentagnostus praecurrens25 sites
- Sidneyia25 sites
The Paleobiology Database records 174 sites for Priapulida, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
77 references- Paleoecology of the Greater Phyllopod Bed community, Burgess Shale81
- Diversity and species abundance patterns of the early Cambrian (Series 2, Stage 3) Chengjiang Biota from China42
- Taphonomy and Palaeoecology of the Early Cambrian Chengjiang Biota from Eastern Yunnan, China40
- Community palaeoecology of the early Cambrian Maotianshan Shale biota: Ecological dominance of priapulid worms17
- Spatial variation in the diversity and composition of the Lower Cambrian (Series 2, Stage 3) Chengjiang Biota, Southwest China15
- Lower Cambrian Burgess Shale-type fossil associations of South China14
- Small shelly fauna from the upper Lower Cambrian Bastion and Ella Island Formations, North-East Greenland13
- A Cambrian soft-bodied biota after the first Phanerozoic mass extinction11
- The Lower Cambrian microfauna of Comley and Rushton, Shropshire, England10
- Palaeoscolecid worms may be nematomorphs rather than annelids9
- The diverse radiodont fauna from the Marjum Formation of Utah, USA (Cambrian: Drumian)7
- Luo et al., 20087
Each of the 386 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 65 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Priapulida
- Source
- Wikipedia/Commons
- Credit
- Shunkina Ksenia
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Priapulida was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Priapulida”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itScalidophora through timeFormations of the CambrianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Théel 1906)