Protoryx
Protoryx is an extinct genus of goat-antelope. A new species, P. tuvaensis, was described by E. L. Dmitrieva and N. V. Serdyuk in 2011, from Russia.

Jonathan Chen · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- BovidaePBDB
- Diet
- grazer, browserPBDB, for Bovidae
- Life habit
- ground dwellingPBDB, for Bovidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Artiodactyla
Time the record covers
At least 8.4 million years on record.
The record covers at least 13.82 Mya to 5.333 Mya. No occurrence read is dated outside 15.98 Mya to 2.58 Mya.
39 occurrences of Protoryx on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
39 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
9 formations
- Mytilinii Formation4
- Kuruksai suite Formation2
- Bahe Formation2
- Vathylakkos Formation2
- Gusar suite Formation1
- Kairubak suite Formation1
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The named rock units Protoryx has been recovered from, largest first. Drawn where the rock is now, not where it formed. 24 occurrences have no formation on record, which is common for older literature.
Protoryx and its relatives
MioceneWhere Protoryx and its relatives were recorded in the Miocene.
Found in the same rock
312 taxa- Gazella15 sites
- Hipparion15 sites
- Choerolophodon pentelici9 sites
- Microstonyx major9 sites
- Prostrepsiceros9 sites
- Ictitherium7 sites
- Palaeoryx7 sites
- Adcrocuta eximia6 sites
- Samotherium boissieri6 sites
- Tragocerus6 sites
- Acerorhinus5 sites
- Chilotherium5 sites
- Machairodus5 sites
- Tragoportax5 sites
- Ancylotherium pentelicum4 sites
- Bovidae4 sites
- Ceratotherium4 sites
- Ctenocervus4 sites
- Deinotherium giganteum4 sites
- Gazella gaudryi4 sites
- Gazella pilgrimi4 sites
- Helicotragus4 sites
- Konobelodon atticus4 sites
- Palaeoreas lindermayeri4 sites
The Paleobiology Database records 29 sites for Protoryx, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
20 references- New excavations in the Neogene mammalian localities of Mytilinii, Samos Island, Greece4
- Upper Cenozoic bio-magnetic stratigraphy of Central Asian mammalian localities4
- Die fossilen Cavicornier von Samos4
- Sur un gisement d’ossements fossiles dans l’île de Samos, contemporains l’âge de Pikermi4
- A hominid tooth from Bulgaria: The last pre-human hominid of continental Europe3
- Mammalian fossils of Samos and Pikermi, Part 1. The Turolian rodents and insectivores of Samos3
- Land mammal faunal sequence of the late Miocene of China: new evidence from Lantian, Shaanxi Province2
- Hipparion from the Bekaa Valley of Lebanon2
- Les vertébrés du Pontien de Salonique2
- The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection 14. Bovidae1
- Bovidae (Mammalia: Artiodactyla) from the Late Miocene of Sivas, Turkey1
- The Neogene mammal localities of Greece: Faunas, chronology and biostratigraphy1
Each of the 39 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 8 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Protoryx
- Source
- Wikipedia/Commons
- Credit
- Jonathan Chen
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Protoryx was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Protoryx”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itBovidae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Forsyth Major 1891