Scarabaeoidea
Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Some of its constituent families are undergoing revision, and the family list below is provisional. This superfamily includes some of the largest beetles extant today, including rhinoceros beetles (Dynastinae), especially the Hercules beetle and the Goliath beetles.

Edmund Reitter · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- scarab beetlePBDB
- Sits inside
- StaphyliniformiaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 199 million years on record.
The record covers at least 199.5 Mya to 11.7 ka. No occurrence read is dated outside 201.4 Mya to 0 Mya.
710 occurrences of Scarabaeoidea on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
710 occurrences- Early Jurassicunder 1%
- Middle Jurassicunder 1%
- Late Jurassic1%
- Early Cretaceous21%
- Late Cretaceous5%
- Paleoceneunder 1%
- Eocene16%
- Oligocene4%
- Miocene16%
- Pliocene2%
- Pleistocene34%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
71 formations
- Highbury Silts and Sands Formation55
- Florissant Formation50
- Yixian Formation · Jehol Group49
- Zaza Formation48
- Shanwang Formation42
- Upper Freshwater-Molasse Formation34
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The named rock units Scarabaeoidea has been recovered from, largest first. Drawn where the rock is now, not where it formed. 47 further formations hold fewer occurrences each and are not listed. 213 occurrences have no formation on record, which is common for older literature.
Scarabaeoidea and its relatives
PleistoceneWhere Scarabaeoidea and its relatives were recorded in the Pleistocene.
Found in the same rock
6,503 taxa- Aphodius64 sites
- Stenus39 sites
- Aleocharinae36 sites
- Carabidae36 sites
- Bembidion29 sites
- Elateridae27 sites
- Coleoptera26 sites
- Curculionidae24 sites
- Helophorus24 sites
- Staphylinidae24 sites
- Chrysomelidae23 sites
- Phyllobius23 sites
- Pterostichus (Cryobius)23 sites
- Morychus viridis21 sites
- Curtonotus alpinus19 sites
- Bledius18 sites
- Donacia18 sites
- Gyrinus18 sites
- Poecilus nearcticus18 sites
- Apion17 sites
- Dytiscidae17 sites
- Hydraena17 sites
- Onthophagus17 sites
- Agabus16 sites
The Paleobiology Database records 305 sites for Scarabaeoidea, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
232 references- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK55
- Late Cenozoic insects of northern Eurasia28
- Zhang, 198928
- Nikolajev, 200722
- Beiträge zur Insektenfauna Oeningens: Coleoptera. Geodephagen, Hydrocanthariden, Gyriniden, Brachelytren, Clavicornen, Lamellicornen und Buprestiden22
- Systematik, Phylogenie, Taphonomie und Paläoökologie der Insekten aus dem Mittel-Eozän des Eckfelder Maares, Vulkaneifel19
- New Miocene Coleoptera from Florissant18
- Heer, 184714
- Mezozoiskie zhestkokryiye13
- Late Pleistocene insects from the Mizozono Formation in Yoshimatsu-cho, Kagoshima Prefecture, Japan12
- Middle Pleistocene deposits at Frog Hall Pit, Stretton-on-Dunsmore, Warwickshire, English Midlands, and their implications for the age of the type Wolstonian12
- Palaeoenvironmental interpretation of Middle and Late Wisconsinan fossil coleopteran assemblages from western Olympic Peninsula, Washington, USA11
Each of the 710 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 220 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Scarabaeoidea
- Source
- Wikipedia/Commons
- Credit
- Edmund Reitter
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Scarabaeoidea was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itStaphyliniformia through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1802