Thylacocephala
Thylacocephala is an extinct group of mandibulate arthropods, that are generally regarded as a kind of crustacean, though their exact position within this group is uncertain. As a class they have a short research history, having been erected in the early 1980s.

Carolin Haug, Derek E G Briggs, Donald G Mikulic, Joanne Kluessendorf, and Joachim T Haug · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- EucrustaceaPBDB
- Diet
- omnivorePBDB
- Life habit
- nektobenthicPBDB
- Motility
- actively mobilePBDB
- Vision
- well-developedPBDB
Time the record covers
At least 371 million years on record.
The record covers at least 457.2 Mya to 85.7 Mya. No occurrence read is dated outside 459.7 Mya to 83.6 Mya.
61 occurrences of Thylacocephala on record, most of them in the Early Triassic. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
61 occurrences- Ordovician3%
- Silurian7%
- Devonian5%
- Carboniferous13%
- Early Triassic31%
- Middle Triassic13%
- Late Triassic7%
- Early Jurassic2%
- Middle Jurassic7%
- Late Jurassic5%
- Early Cretaceous2%
- Late Cretaceous6%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
20 formations
- Nanlinghu Formation7
- Buchenstein Formation7
- Osawa Formation5
- Swope Formation · Kansas City Group3
- Brandon Bridge Formation3
- Carbondale Formation · Kewanee Group3
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The named rock units Thylacocephala has been recovered from, largest first. Drawn where the rock is now, not where it formed. 15 occurrences have no formation on record, which is common for older literature.
Thylacocephala and its relatives
Early TriassicWhere Thylacocephala and its relatives were recorded in the Early Triassic.
Found in the same rock
414 taxa- Concavicaris sinuata4 sites
- Gorgonophontes peleron4 sites
- Mycterops whitei4 sites
- Orbiculoidea4 sites
- Aeger3 sites
- Angustidontus seriatus3 sites
- Ankitokazocaris lariensis3 sites
- Bivalvia3 sites
- Bobasatrania3 sites
- Ligulacaris parisiana3 sites
- Microcaris minuta3 sites
- Parisicaris triassica3 sites
- Penaeoidea3 sites
- Aeschronectida2 sites
- Ammonoidea indet.2 sites
- Ankitokazocaris2 sites
- Ankitokazocaris chaohuensis2 sites
- Ankitokazocaris utatsuensis2 sites
- Brachyphyllum2 sites
- Ceratiocaris papilio2 sites
- Coelacanthidae2 sites
- Coleoidea2 sites
- Concavicaris2 sites
- Conodonta2 sites
The Paleobiology Database records 38 sites for Thylacocephala, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
27 references- New material of Thylacocephala from the early Ladinian (Middle Triassic) of Northern Grigna (Lecco, Lombardy, Northern Italy)7
- New Thylacocephala (Crustacea) assemblage from the Spathian (Lower Triassic) of Majiashan (Chaohu, Anhui Province, South China)6
- Exceptional fossil assemblages confirm the existence of complex Early Triassic ecosystems during the early Spathian4
- Thylacocephala (Arthropoda) from the Lower Triassic of the South Kitakami Belt, northeast Japan4
- Remarkable Invertebrates from the Lower Callovian of La Voulte-sur-Rhône (Ardèche, France)4
- Upper Pennsylvanian arthropods from black shales of Iowa and Nebraska4
- New species of Thylacocephala, Eodollocaris keithflinti n. gen., n. sp., from the Mazon Creek Lagerstätte, Illinois, United States (c. 307 Ma) and redescription of other Mazon Creek thylacocephalans3
- New thylacocephalans from the Early Triassic Paris Biota (Bear Lake County, Idaho, USA)3
- The implications of a Silurian and other thylacocephalan crustaceans for the functional morphology and systematic affinities of the group3
- Upper Ordovician Thylacocephala (Euarthropoda, Eucrustacea) from Bohemia indicate early ecological differentiation2
- First Mesozoic thylacocephalans (Arthropoda, Crustacea; Cretaceous) in the western hemisphere: new discoveries from the Muhi Quarry Lagerstätte2
- Angustidontus, a Late Devonian pelagic predatory crustacean2
Each of the 61 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 15 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Thylacocephala
- Source
- Wikipedia/Commons
- Credit
- Carolin Haug, Derek E G Briggs, Donald G Mikulic, Joanne Kluessendorf, and Joachim T Haug
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Thylacocephala was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itEucrustacea through timeFormations of the Early TriassicThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Pinna et al. 1982