Tragelaphus angasii
The lowland nyala or simply nyala is a spiral-horned artiodactyl antelope native to Southern Africa. The species is part of the family Bovidae and the genus Tragelaphus. It was first described in 1849 by George French Angas and exhibits the highest sexual dimorphism among the spiral-horned antelopes. It is not to be confused with the endangered mountain nyala living in the Bale region of Ethiopia.

Charles J. Sharp · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- nyalaPBDB
- Sits inside
- TragelaphusPBDB
- Diet
- browserPBDB, for Tragelaphus
- Life habit
- ground dwellingPBDB, for Bovidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Artiodactyla
Time the record covers
At most 24 million years on record.
Every occurrence read is dated to somewhere between 23.04 Mya and 0 Mya, and nothing in the dating separates them, so that window is what the record shows rather than a span.
8 occurrences of Tragelaphus angasii on record, most of them in the Pliocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
8 occurrences- Miocene16%
- Pliocene42%
- Pleistocene42%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
6 formations
- Chiwondo Formation2
- Varswater & Bredasdorp Formation1
- Makapansgat Formation1
- Sterkfontein Formation · Transvaal Group1
- Swartkrans Formation · Transvaal Group1
- Varswater Formation1
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The named rock units Tragelaphus angasii has been recovered from, largest first. Drawn where the rock is now, not where it formed. 1 occurrence has no formation on record, which is common for older literature.
Tragelaphus angasii and its relatives
PlioceneWhere Tragelaphus angasii and its relatives were recorded in the Pliocene.
Found in the same rock
273 taxa- Equus5 sites
- Gazella4 sites
- Hipparion4 sites
- Tragelaphus strepsiceros4 sites
- Bovidae3 sites
- Canidae3 sites
- Caracal caracal3 sites
- Cercopithecidae3 sites
- Damaliscus3 sites
- Diceros bicornis3 sites
- Elephantulus3 sites
- Equus capensis3 sites
- Hyaenidae3 sites
- Hylochoerus euilus3 sites
- Hystrix africaeaustralis3 sites
- Lupulella mesomelas3 sites
- Palaeoloxodon recki3 sites
- Panthera pardus3 sites
- Phenacotragus vanhoepeni3 sites
- Procavia antiqua3 sites
- Suidae3 sites
- Suncus3 sites
- Taurotragus oryx3 sites
- Adenota recki2 sites
The Paleobiology Database records 8 sites for Tragelaphus angasii, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
7 references- Paleobiogeography of the Malawi Rift: Age and vertebrate paleontology of the Chiwondo Beds, northern Malawi2
- Paleoecology of the late Tertiary fossil occurences in 'E' Quarry, Langebaanweg, South Africa and a reinterpretation of their geological context1
- Re-appraisal of lithostratigraphy of Sterkfontein hominid site1
- The Mammalian Fauna from the Middle and Later Stone Age (Later Pleistocene) Levels of Border Cave, Natal Province, South Africa1
- Re-interpretation of the Swartkrans site and its remains1
- Elephant Shrews (Macroscelididae) from Olduvai and Makapansgat1
- The Pliocene fossil occurences in 'E' Quarry, Langebaanweg, South Africa1
Each of the 8 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Tragelaphus angasii
- Source
- Wikipedia/Commons
- Credit
- Charles J. Sharp
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Tragelaphus angasii was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Nyala”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itTragelaphus through timeFormations of the PlioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Gray 1849