Ugandax
Ugandax is an extinct genus of bovines in the subtribe Bubalina that lived from the Miocene to the Pleistocene of Africa. Cladistic analyses suggest Ugandax represents an ancestral form of the African buffalo, Syncerus, and teeth assigned to Ugandax represent the earliest appearance of bovines in Africa.

Apokryltaros · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- BoviniPBDB
- Diet
- grazer, browserPBDB, for Bovidae
- Life habit
- ground dwellingPBDB, for Bovidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Artiodactyla
Time the record covers
At least 2.7 million years on record.
The record covers at least 5.333 Mya to 2.58 Mya. No occurrence read is dated outside 23.04 Mya to 11.7 ka.
29 occurrences of Ugandax on record, most of them in the Pliocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
29 occurrences- Miocene33%
- Pliocene42%
- Pleistocene26%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
9 formations
- Nachukui Formation · Omo Group7
- Nkondo Formation · Albertine Group6
- Adu-Asa Formation · Awash Group3
- Hadar Formation · Awash Group3
- Varswater Formation3
- Chiwondo Formation2
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The named rock units Ugandax has been recovered from, largest first. Drawn where the rock is now, not where it formed. 2 occurrences have no formation on record, which is common for older literature.
Ugandax and its relatives
PlioceneWhere Ugandax and its relatives were recorded in the Pliocene.
Found in the same rock
329 taxa- Hipparion13 sites
- Kobus13 sites
- Gazella11 sites
- Giraffa10 sites
- Hexaprotodon10 sites
- Palaeoloxodon recki10 sites
- Alcelaphini9 sites
- Homotherium9 sites
- Deinotherium bozasi8 sites
- Tragelaphus nakuae8 sites
- Ceratotherium7 sites
- Hylochoerus euilus7 sites
- Metridiochoerus andrewsi7 sites
- Reduncini7 sites
- Tragelaphus7 sites
- Aepyceros shungurae6 sites
- Cercopithecidae6 sites
- Gerontochoerus scotti6 sites
- Nyanzachoerus kanamensis6 sites
- Suidae6 sites
- Aepyceros5 sites
- Diceros bicornis5 sites
- Hippotragini5 sites
- Libytherium maurusium5 sites
The Paleobiology Database records 28 sites for Ugandax, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
18 references- Stratigraphy and Paleontology of pliocene and Pleistocene Localities West of Lake Turkana, Kenya7
- Evolution of bovid diversity in the Plio-Pleistocene of Africa4
- Refined stratigraphy of the hominid-bearing Awash Group, Middle Awash Valley, Afar Depression, Ethiopia2
- Paleontological excavations in the Shungura Formation, Lower Omo Basin, 1969-19732
- A fossil terrestrial fauna from Tobène (Senegal) provides a unique early Pliocene window in western Africa1
- Mio-Pliocene Mammals from the Middle Awash, Ethiopia1
- Extinct ostrich eggshell (Aves: Struthionidae) from the Pliocene Chiwondo Beds, Malawi: implications for the potential biostratigraphic correlation of African Neogene deposits1
- Cainozoic mammals from coastal Namaqualand, South Africa1
- Paleobiogeography of the Malawi Rift: Age and vertebrate paleontology of the Chiwondo Beds, northern Malawi1
- Neogene and Pleistocene squamates from the Western Rift,Uganda1
- Equidae of the Albertine rift valley,Uganda1
- Carnivora from Nkondo and Kisegi-Nyabusosi,Western Rift, Uganda1
Each of the 29 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 6 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ugandax
- Source
- Wikipedia/Commons
- Credit
- Apokryltaros
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Ugandax was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Ugandax”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itBovini through timeFormations of the PlioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Cooke and Coryndon 1970