Anomala
Anomala is a genus of shining leaf chafers in the scarab beetle family Scarabaeidae. There are more than 1,100 described species in Anomala, found in every continent except Antarctica.

Visviva · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- AnomalinaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 33 million years on record.
The record covers at least 33.9 Mya to 774 ka. No occurrence read is dated outside 37 Mya to 129 ka.
28 occurrences of Anomala on record, most of them in the Miocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
28 occurrences- Eocene7%
- Oligocene18%
- Miocene68%
- Pliocene4%
- Pleistocene4%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
6 formations
- Shanwang Formation18
- Rott Formation3
- Florissant Formation2
- Upper Freshwater-Molasse Formation1
- Highbury Silts and Sands Formation1
- Shiake Formation1
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The named rock units Anomala has been recovered from, largest first. Drawn where the rock is now, not where it formed. 2 occurrences have no formation on record, which is common for older literature.
Anomala and its relatives
MioceneWhere Anomala and its relatives were recorded in the Miocene.
Found in the same rock
916 taxa- Limois shanwangensis5 sites
- Calosoma maderae4 sites
- Camponotites shanwangensis4 sites
- Cerambycidae4 sites
- Dascillus shandongianus4 sites
- Gerstaeckerus alviolatus4 sites
- Hylobius plenus4 sites
- Bibio shanwangensis3 sites
- Chlaenius furvus3 sites
- Curculionidae3 sites
- Entiminae3 sites
- Epiaeschna lucida3 sites
- Formica3 sites
- Heptagenia3 sites
- Scarabaeoidea3 sites
- Abia shandongensis2 sites
- Allodahlia shanwangensis2 sites
- Anisoptera2 sites
- Aphaena lithoecia2 sites
- Apidae2 sites
- Apioninae2 sites
- Apis (Synapis) miocenica2 sites
- Atactogaster shanwangensis2 sites
- Bibio cratopodus2 sites
The Paleobiology Database records 13 sites for Anomala, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
11 references- Zhang, 198911
- Zhang et al., 19945
- Käfer und Polypen aus der Braunkohle des Siebengebirges3
- New Miocene Coleoptera from Florissant2
- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK1
- Miocene Coleoptera (Insecta) of Shanwang, Shandong Province, China1
- Fossil insects in the diatoms of Shanwang1
- Late Cenozoic insect faunas in the northwestern region of Kagoshima Prefecture, Japan1
- Poissons, crustacés et insectes fossiles de l'Oligocène du Puy-de-Mur (Auvergne)1
- Quelques insectes de l'Aquitanien de Rott, Sept-Monts (Prusse rhénane)1
- Beiträge zur Insektenfauna Oeningens: Coleoptera. Geodephagen, Hydrocanthariden, Gyriniden, Brachelytren, Clavicornen, Lamellicornen und Buprestiden1
Each of the 28 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Anomala
- Source
- Wikipedia/Commons
- Credit
- Visviva
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Anomala was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Anomala”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAnomalina through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Samouelle 1819