Caenophidia
The Caenophidia are a derived clade of alethinophidian snakes, which contains over 80% of all the extant species of snakes. The largest family is Colubridae, but it also includes at least seven other families, at least four of which were once classified as "Colubridae" before molecular phylogenetics helped us understand their relationships. It has been found to be monophyletic.

Clinton & Charles Robertson from Del Rio, Texas & San Marcos, TX, USA · CC BY 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- AlethinophidiaPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 60 million years on record.
The record covers at least 66 Mya to 5.09 Mya. No occurrence read is dated outside 83.6 Mya to 3.55 Mya.
26 occurrences of Caenophidia on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
26 occurrences- Late Cretaceous8%
- Paleocene1%
- Eocene45%
- Miocene30%
- Pliocene16%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
9 formations
- Komnina Formation3
- Wadi Milk Formation2
- Cambay Shale Formation2
- Birket Qarun Formation2
- Prolemais Formation2
- Mansar Formation · Siwalik Group1
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The named rock units Caenophidia has been recovered from, largest first. Drawn where the rock is now, not where it formed. 11 occurrences have no formation on record, which is common for older literature.
Caenophidia and its relatives
EoceneWhere Caenophidia and its relatives were recorded in the Eocene.
Found in the same rock
502 taxa- Anura8 sites
- Russellophiidae7 sites
- Serpentes7 sites
- Latonia6 sites
- Ranidae6 sites
- Apodemus5 sites
- Boidae5 sites
- Acrochordus dehmi4 sites
- Anguinae4 sites
- Anguis4 sites
- Crocodylia4 sites
- Lacertidae4 sites
- Micromys bendai4 sites
- Natricinae4 sites
- Rana4 sites
- Testudinoidea4 sites
- Trionyx4 sites
- Anguidae3 sites
- Anguini3 sites
- Apodemus atavus3 sites
- Batagur3 sites
- Brachypotherium3 sites
- Channa3 sites
- Claridae indet.3 sites
The Paleobiology Database records 23 sites for Caenophidia, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
15 references- Amphibians and reptiles from the late Miocene and early Pliocene of the Ptolemais area (Western Macedonia, Greece)6
- Miocene vertebrates from the Siwalik Group, Western Nepal3
- A late Eocene snake fauna from the Fayum Depression, Egypt2
- A diverse snake fauna from the early Eocene of Vastan Lignite Mine, Gujarat, India2
- Late Eocene snakes from the Headon Hill Formation, southern England2
- Les amphibiens et les reptiles de Prémontré (Cuisien, Bassin parisien): une herpétofaune de référence pour l'Eocène inférieur2
- A Middle Miocene (~14 Ma) vertebrate assemblage from Palasava, Rapar Taluka, Kutch (Kachchh) District, Gujarat State, western India1
- Squamate reptiles from the middle Eocene of Lissieu (France). A landmark in the middle Eocene of Europe1
- Fossil snakes from the Palaeocene of São José de Itaboraí, Brazil. Part III. Ungaliophiinae, booids incertae sedis, and Caenophidia. Summary, update and discussion of the snake fauna from the locality1
- Amphibians and squamates from the Neogene Siwalik beds of Jammu and Kashmir, India1
- Mid-Cretaceous (Cenomanian) snakes from Wadi Abu Hashim, Sudan: The earliest snake assemblage.1
- A dinosaur fauna from the Late Cretaceous (Cenomanian) of northern Sudan1
Each of the 26 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 3 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Caenophidia
- Source
- Wikipedia/Commons
- Credit
- Clinton & Charles Robertson from Del Rio, Texas & San Marcos, TX, USA
- Licence
- CC BY 2.0
- Commercial use
- the licence allows it
No 3D scan of Caenophidia was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Caenophidia”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAlethinophidia through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Hoffstetter 1939)