Chlaenius
Chlaenius is a large and diverse genus of ground beetle. It is native to the Palearctic realm, Afrotropical realm, and Nearctic realm. Worldwide, roughly 1,000 species are currently recognized with most known species occurring in the Oriental and Afrotropical regions. The genus is divided into many subgenera.

Edmund Reitter · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ChlaeniinaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 23 million years on record.
The record covers at least 23.04 Mya to 11.7 ka. No occurrence read is dated outside 27.3 Mya to 0 Mya.
20 occurrences of Chlaenius on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
20 occurrences- Oligocene5%
- Miocene25%
- Pleistocene65%
- Holocene5%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
8 formations
- Shanwang Formation5
- Ookui Formation2
- Tsumori Formation2
- Highbury Silts and Sands Formation2
- Kap København Formation1
- Nogami Formation1
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The named rock units Chlaenius has been recovered from, largest first. Drawn where the rock is now, not where it formed. 5 occurrences have no formation on record, which is common for older literature.
Chlaenius and its relatives
PleistoceneWhere Chlaenius and its relatives were recorded in the Pleistocene.
Found in the same rock
818 taxa- Curculionidae9 sites
- Agabus7 sites
- Carabidae7 sites
- Coleoptera6 sites
- Aphodius5 sites
- Staphylinidae5 sites
- Bembidion4 sites
- Carabus4 sites
- Cerambycidae4 sites
- Donacia4 sites
- Donaciinae4 sites
- Elateridae4 sites
- Gyrinus4 sites
- Helophorus4 sites
- Lepidoptera4 sites
- Pterostichus4 sites
- Stenus4 sites
- Aleocharinae3 sites
- Calosoma maderae3 sites
- Camponotites shanwangensis3 sites
- Carpelimus3 sites
- Chaetocnema3 sites
- Coccinellidae3 sites
- Coelostoma orbiculare3 sites
The Paleobiology Database records 17 sites for Chlaenius, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
15 references- Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK2
- Middle Pleistocene fossil insects from the Tsumori Formation of Mashiki-machi,Kumamoto Prefecture,Japan (Part 2)2
- Early Pleistocene ground beetles (Coleoptera: Carabidae) from the Ookui Formation in Nagano Prefecture, central Japan, and their biogeographical and paleoenvironmental significance2
- Zhang, 19892
- Miocene Coleoptera (Insecta) of Shanwang, Shandong Province, China2
- The paleoenvironment of the Olympia beds based on fossil beetles from Discovery Park, Seattle, Washington, U.S.A.1
- Middle Pleistocene fossil insects from the Nogami Formation of Kokonoe, Oita Prefecture, Kyushu, Japan1
- Early Pleistocene fossil insects from the Oyama Formation at Oyama-machi, Oita Prefecture, Kyushu, Japan1
- Paleogeography of the lower Volga at the end of Atelian time based on entomological and botanical data1
- Palaeoentomology of the Kap København Formation, a Plio-Pleistocene sequence in Peary Land, North Greenland1
- Fossil insects in the diatoms of Shanwang1
- Poissons, crustacés et insectes fossiles de l'Oligocène du Puy-de-Mur (Auvergne)1
Each of the 20 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 3 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Chlaenius
- Source
- Wikipedia/Commons
- Credit
- Edmund Reitter
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Chlaenius”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itChlaeniina through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Bonelli 1810