Ischyodus
Ischyodus is an extinct genus of chimaera. It is the most diverse and long-lived chimaera genus, with over 39 species found worldwide spanning over 140 million years from the Middle Jurassic to the Miocene. Almost all species are only known from tooth plates, with the exception of the Jurassic species I. quenstedti. Complete specimens of I. quenstedti from the Late Jurassic of Germany most closely resemble the genus Callorhinchus amongst living chimaera genera. It is sometimes placed in the "Edaphodontidae", a unclearly defined group of chimaera with an uncertain position within the clade, while other authors place it into Callorhinchidae along with Callorhinchus, a position that has been supported by at least some phylogenetic analyses. It is suggested therefore to probably have had a similar ecology to living Callorhinchus.

Ghedoghedo · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- CallorhynchidaePBDB
- Diet
- carnivorePBDB, for Chimaeriformes
- Life habit
- nektobenthicPBDB, for Chimaeriformes
- Motility
- actively mobilePBDB, for Chimaeriformes
Time the record covers
At least 159 million years on record.
The record covers at least 165.3 Mya to 5.333 Mya. No occurrence read is dated outside 168.2 Mya to 3.6 Mya.
94 occurrences of Ischyodus on record, most of them in the Late Cretaceous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
94 occurrences- Middle Jurassic5%
- Late Jurassic9%
- Early Cretaceous5%
- Late Cretaceous50%
- Paleocene12%
- Eocene14%
- Miocene3%
- Pliocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
45 formations
- Hell Creek Formation · Montana Group9
- La Meseta Formation9
- Fox Hills Formation · Montana Group5
- Hornerstown Formation · Rancocas Group4
- Mount Laurel Formation · Monmouth Group3
- Judith River Formation · Montana Group3
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The named rock units Ischyodus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 21 further formations hold fewer occurrences each and are not listed. 13 occurrences have no formation on record, which is common for older literature.
Ischyodus and its relatives
Late CretaceousWhere Ischyodus and its relatives were recorded in the Late Cretaceous.
Found in the same rock
978 taxa- Squalicorax kaupi18 sites
- Ischyrhiza mira17 sites
- Archaeolamna kopingensis15 sites
- Cretalamna appendiculata15 sites
- Scapanorhynchus texanus13 sites
- Squalicorax pristodontus13 sites
- Carcharias samhammeri11 sites
- Hybodus11 sites
- Paralbula casei11 sites
- Squatina11 sites
- Squatina (Cretascyllium) hassei11 sites
- Anomoeodus phaseolus10 sites
- Brachyrhizodus wichitaensis10 sites
- Enchodus ferox10 sites
- Myliobatis10 sites
- Odontaspis aculeatus10 sites
- Rhombodus levis10 sites
- Striatolamia macrota10 sites
- Crocodylia9 sites
- Paranomotodon angustidens9 sites
- Carcharias holmdelensis8 sites
- Hadrosauridae8 sites
- Enchodus7 sites
- Palaeohypotodus rutoti7 sites
The Paleobiology Database records 88 sites for Ischyodus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
59 references- A new species of Ischyodus (Chondrichthyes: Holocephali: Callorhynchidae) from upper Maastrichtian shallow marine facies of the Fox Hills and Hell Creek formations, Williston Basin, North Dakota, USA13
- Paleocene chimaeroid fishes (Chondrichthyes: Holocephali) from the eastern United States, including two new species of Callorhinchus5
- Chimaeroid fish remains from Seymour Island, Antarctic Peninsula5
- Middle Eocene chondrichthyan fauna from Antarctic Peninsula housed in the Museo de La Plata, Argentina3
- A review of the Tertiary fossil Cetacea (Mammalia) localities in Australia3
- Paleoecology of the Delaware Valley region. Part II: Cretaceous to Quartenary3
- Catalogue of the Fossil Fishes in the British Museum (Natural History). Part II3
- The fossil record of chimaeras (Chondrichthyes, Holocephali) in Argentina2
- Ultimate Eocene (Priabonian) chondrichthyans (Holocephali, Elasmobranchii) of Antarctica2
- An embryonic mandibular tooth plate and associated remains of a Late Jurassic chimaeroid (Holocephali, Chimaeriformes) from the Iberian Peninsula2
- Additional fish records from the Judith River Formation (Campanian) of Montana2
- Ischyodus bifurcatus, a new species of chimaeroid fish from the Upper Cretaceous of New Jersey2
Each of the 94 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 47 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ischyodus
- Source
- Wikipedia/Commons
- Credit
- Ghedoghedo
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Ischyodus”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCallorhynchidae through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Egerton 1843