Miliolida
The Miliolida are an order of foraminifera with calcareous, porcelacous tests that are imperforate and commonly have a pseudochitinous lining. Tests are composed of randomly oriented calcite needles that have a high proportion of magnesium along with organic material. Tests lack pores and generally have multiple chambers.

Wilson44691 · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- TubothalameaPBDB
- Diet
- omnivorePBDB, for Foraminifera
- Life habit
- semi-infaunalPBDB, for Foraminifera
- Motility
- stationaryPBDB, for Foraminifera
Time the record covers
At least 351 million years on record.
The record covers at least 351.9 Mya to 11.7 ka. No occurrence read is dated outside 358.86 Mya to 0 Mya.
8,613 occurrences of Miliolida on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
8,613 occurrences- Carboniferous2%
- Permian9%
- Early Triassic2%
- Middle Triassic2%
- Late Triassic4%
- Early Jurassicunder 1%
- Middle Jurassicunder 1%
- Late Jurassic1%
- Early Cretaceous3%
- Late Cretaceous7%
- Paleocene5%
- Eocene8%
- Oligocene6%
- Miocene11%
- Pliocene6%
- Pleistocene30%
- Holocene1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
437 formations
- Qom Formation Formation380
- Polat Formation · Geyik Dagi Unit Group180
- Sylhet Formation Formation163
- Sinjar Formation99
- Wujiaping Formation74
- Oread Limestone Formation · Shawnee Group69
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The named rock units Miliolida has been recovered from, largest first. Drawn where the rock is now, not where it formed. 413 further formations hold fewer occurrences each and are not listed. 5,592 occurrences have no formation on record, which is common for older literature.
Miliolida and its relatives
PleistoceneWhere Miliolida and its relatives were recorded in the Pleistocene.
Found in the same rock
3,148 taxa- Nubeculinella46 sites
- Serpula46 sites
- Bryozoa42 sites
- Nanogyra (Nanogyra) nana41 sites
- Gastropoda37 sites
- Ophthalmidium37 sites
- Actinostreon gregarea31 sites
- Quinqueloculina28 sites
- Glomospira26 sites
- Echinoidea25 sites
- Gryphaea25 sites
- Nodosaria25 sites
- Ammobaculites24 sites
- Bivalvia24 sites
- Ostracoda23 sites
- Shamovella obscura22 sites
- Isognomon21 sites
- Tetrataxis21 sites
- Miliolidae20 sites
- Radulopecten fibrosus20 sites
- Textularia20 sites
- Tubiphytes20 sites
- Endothyra19 sites
- Galeanella18 sites
The Paleobiology Database records at least 1,054 sites for Miliolida, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
700 references- Sedimentary facies and depositional environments of the Oligocene–early Miocene marine Qom Formation, Central Iran Back-Arc Basin, Iran (northeastern margin of the Tethyan Seaway)380
- Early and Middle Triassic trends in diversity, evenness, and size of foraminifers on a carbonate platform in south China: Implications for tempo and mode of biotic recovery from the end-Permian mass extinction265
- Taxonomic and Paelogeographic Approaches to the Dasyclad Algae in the Upper Jurassic (Kimmeridgian)-Upper Cretaceous (Cenomanian) Peritidal Carbonates of the Fele (YassÝbel) Area (Western Taurides, Turkey)180
- The Enigmatic Paleocene-Eocene coralline Distichoplax: Approaching the structural complexities, ecological affinities and extinction hypotheses163
- Site U1337157
- Depositional History of a Paleogene Carbonate Ramp (Western Cephalonia, Ionian Islands, Greece)145
- Site U1352128
- Site U1482126
- Site U1386126
- Site U1391116
- Development of rudist lithosomes in the Coniacian-Lower Campanian carbonate shelves of central-southern Italy: high-energy vs low-energy settings105
- Site U1354104
Each of the 8,613 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 688 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Miliolida
- Source
- Wikipedia/Commons
- Credit
- Wilson44691
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan of Miliolida was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Miliolida”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itTubothalamea through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Lankester 1885