Zanthoxylum
Zanthoxylum is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae in the subfamily Rutoideae. Several of the species have yellow heartwood, to which their generic name alludes. Several species are cultivated for their use as spices, notably including Sichuan pepper.

Forest & Kim Starr · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- prickly ashPBDB
- Sits inside
- RutaceaePBDB
- Diet
- "photoautotroph"PBDB, for Angiospermae
Time the record covers
At least 45 million years on record.
The record covers at least 48.07 Mya to 2.58 Mya. No occurrence read is dated outside 56 Mya to 11.7 ka.
29 occurrences of Zanthoxylum on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
29 occurrences- Eocene17%
- Oligocene14%
- Miocene48%
- Pliocene11%
- Pleistocene10%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
10 formations
- Headon Beds Formation2
- San Sebastian Formation · Rio Guatemala Group2
- Sanying Formation2
- Ngorora Formation1
- Oldhaven Formation1
- Bangmai Formation1
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The named rock units Zanthoxylum has been recovered from, largest first. Drawn where the rock is now, not where it formed. 16 occurrences have no formation on record, which is common for older literature.
Zanthoxylum and its relatives
MioceneWhere Zanthoxylum and its relatives were recorded in the Miocene.
Found in the same rock
602 taxa- Celtis6 sites
- Cyathea5 sites
- Ficus5 sites
- Alnus4 sites
- Arecaceae4 sites
- Engelhardia4 sites
- Mallotus4 sites
- Meliosma4 sites
- Podocarpus4 sites
- Pteris4 sites
- Quercus4 sites
- Symplocos4 sites
- Syzygium4 sites
- Acacia3 sites
- Acalypha3 sites
- Acer3 sites
- Agathis3 sites
- Aleurites3 sites
- Alphitonia3 sites
- Ampelopsis3 sites
- Araliaceae3 sites
- Araucaria3 sites
- Archontophoenix3 sites
- Argyrodendron3 sites
The Paleobiology Database records 17 sites for Zanthoxylum, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
14 references- Die Miozaenan Frucht-und Samen-Floren die Oberpfaelzer Braunkohle. I. Funde aus den sandigen zwischenmitteln7
- Continuous existence of Zanthoxylum (Rutaceae) in Southwest China since the Miocene3
- A complete pollen record of the last 230 ka from Lynch's Crater, north-eastern Australia3
- A catalogue of Cenozoic megafossil plants in China3
- Die Floren des Haselbacher Serie im Weisselster-Becken (Bezirk Leipzig, DDR)2
- Studies in neotropical paleobotany. I. The Oligocene communities of Puerto Rico2
- Flora of the Lower Headon Beds of Hampshire and the Isle of Wight2
- Quantitative reconstruction of the Late Miocene monsoon climates of southwest China: A case study of the Lincang flora from Yunnan Province1
- Angiospermous fossil leaves from the Siwalik foreland basins and their palaeoclimatic implications1
- Late Eocene silicified fruits and seeds from the John Day Formation near Post, Oregon1
- Re-Evaluation of the age of the Brandon lignite (Vermont, USA) based on plant megafossils1
- A middle Miocene (12.2 my old) forest in the East African Rift Valley, Keny1
Each of the 29 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 2 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Zanthoxylum
- Source
- Wikipedia/Commons
- Credit
- Forest & Kim Starr
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Zanthoxylum was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Zanthoxylum”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itRutaceae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Linnaeus 1753