Testudines
Searched as Chelonia. The Paleobiology Database treats Testudines as the accepted name.

Ernst Haeckel · Public Domain · via Wikidata/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- turtlePBDB
- Sits inside
- TestudinataPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Diapsida
Time the record covers
At least 246 million years on record.
The record covers at least 247 Mya to 11.7 ka. No occurrence read is dated outside 250.8 Mya to 0 Mya.
9,209 occurrences of Testudines on record, most of them in the Late Cretaceous. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
9,209 occurrences- Early Triassicunder 1%
- Middle Triassicunder 1%
- Late Triassicunder 1%
- Early Jurassicunder 1%
- Middle Jurassicunder 1%
- Late Jurassic1%
- Early Cretaceous4%
- Late Cretaceous29%
- Paleocene6%
- Eocene17%
- Oligocene4%
- Miocene19%
- Pliocene5%
- Pleistocene11%
- Holocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
1194 formations
- Hell Creek Formation · Montana Group350
- Denver Formation156
- Willwood Formation150
- Kirtland Formation149
- Bridger Formation124
- Uinta Formation120
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The named rock units Testudines has been recovered from, largest first. Drawn where the rock is now, not where it formed. 1,170 further formations hold fewer occurrences each and are not listed. 2,704 occurrences have no formation on record, which is common for older literature.
Testudines and its relatives
Late CretaceousWhere Testudines and its relatives were recorded in the Late Cretaceous.
Found in the same rock
3,332 taxa- Trionychidae46 sites
- Geochelone40 sites
- Bovidae37 sites
- Crocodylus35 sites
- Trionyx35 sites
- Clarias32 sites
- Hipparion30 sites
- Lepisosteus28 sites
- Rhinocerotidae28 sites
- Ceratotherium27 sites
- Gazella27 sites
- Equus26 sites
- Aves25 sites
- Crocodylus niloticus24 sites
- Euthecodon brumpti24 sites
- Crocodylia23 sites
- Cercopithecidae22 sites
- Lates21 sites
- Mammalia21 sites
- Polypterus21 sites
- Testudinidae21 sites
- Alcelaphini20 sites
- Hominidae20 sites
- Osteichthyes20 sites
The Paleobiology Database records at least 1,144 sites for Testudines, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
2,626 references- Contribution a l'etude des Cheloniens. Cheloniens continentaux due Cretace et du Tertiare de France251
- Re-assessment of Late Campanian (Kirtlandian) turtles from the Upper Cretaceous Fruitland and Kirtland formations, San Juan Basin, New Mexico, USA191
- Patterns of geographic variation in latest Cretaceous vertebrates: evidence from the turtle component157
- Systématique du groupe "Palaeochelys sensu lato – Mauremys" (Chelonii, Testudinoidea) du Tertiaire d'Europe occidentale: principaux résultats [Systematic of the "Palaeochelys sensu lato – Mauremys" group (Chelonii, Testudinoidea) from the Tertiary of Western Europe: principal results107
- Vertebrate biostratigraphy of the Hell Creek Formation in southwestern North Dakota and northwestern South Dakota107
- Fossil Vertebrate Collections104
- Exceptional continental record of biotic recovery after the Cretaceous–Paleogene mass extinction94
- The fossil turtles of North America93
- Contribution to the diversity of the fossil record of turtles (Testudinidata) from Chubut province (Argentina) and its significance in understanding the evolution of turtles in southern South America88
- New Asiatic materials on turtles of the family Adocidae with a review of the adocid record in Asia72
- Synonymies and reidentifications of North American fossil vertebrates and so forth69
- New turtles from the Paleogene of North America67
Each of the 9,209 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 2,614 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3

Picture of Testudines
- Source
- Wikidata/Commons
- Credit
- Ernst Haeckel
- Licence
- Public Domain
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Chelonia mydas · smf:63250
3D model
- Source
- Sketchfab
- Licence
- CC Attribution-NonCommercial
- Commercial use
- the licence does not allow it
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itTestudinata through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Latreille 1800)