Gymnura
The butterfly rays are the rays forming the genus Gymnura and the family Gymnuridae. They are found in warm oceans worldwide, and occasionally in estuaries.

Tomas Willems & Hans Hillewaert · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- GymnuridaePBDB
- Diet
- carnivorePBDB, for Myliobatiformes
- Life habit
- nektobenthicPBDB, for Myliobatiformes
- Motility
- actively mobilePBDB, for Myliobatiformes
Time the record covers
At least 88 million years on record.
The record covers at least 93.9 Mya to 5.333 Mya. No occurrence read is dated outside 100.5 Mya to 3.6 Mya.
46 occurrences of Gymnura on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
46 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
25 formations
- Birket Qarun Formation4
- Selsey Formation · Bracklesham Group3
- Gehannam Formation3
- Marnes de Miretrain Formation3
- Esbarrondadoiro Formation3
- Couche II Formation3
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The named rock units Gymnura has been recovered from, largest first. Drawn where the rock is now, not where it formed. One further formation holds fewer occurrences and is not listed. 4 occurrences have no formation on record, which is common for older literature.
Gymnura and its relatives
EoceneWhere Gymnura and its relatives were recorded in the Eocene.
Found in the same rock
677 taxa- Dasyatis20 sites
- Rhinobatos12 sites
- Scyliorhinus12 sites
- Heterodontus vincenti11 sites
- Mustelus11 sites
- Myliobatis11 sites
- Galeorhinus9 sites
- Isurus praecursor9 sites
- Abdounia8 sites
- Burnhamia daviesi8 sites
- Carcharhinus8 sites
- Coupatezia8 sites
- Isurolamna inflata8 sites
- Physogaleus8 sites
- Rhizoprionodon8 sites
- Carcharias7 sites
- Coupatezia woutersi7 sites
- Pararhincodon germaini7 sites
- Premontreia (Oxyscyllium) gilberti7 sites
- Rhinoptera7 sites
- Anoxypristis6 sites
- Burnhamia6 sites
- Dasyatidae6 sites
- Hemiscyllium tailledisensis6 sites
The Paleobiology Database records 38 sites for Gymnura, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
25 references- Shark and ray faunas in the Middle and Late Eocene of the Fayum Area, Egypt8
- Nouvelle faunes de sélaciens (Elasmobranchii, Neoselachii) de l'Eocène moyen des Landes (Sud-Ouest, France). Implication dans la connaissance des communautés de sélaciens d'eau profondes6
- Les Orectolobiformes, Carcharhiniformes et Myliobatiformes (Elasmobranchii, Neoselachii) des bassins à phosphate du Maroc (Maastrichtien-Lutétien basal). Systématique, biostratigraphie, évolution et dynamique des faunes4
- Sélaciens du Miocène terminal du bassin d'Alvalade (Portugal) Essai de synthèse3
- The Selsey Division (Bracklesham Group) at Lee-on-the-Solent, Gosport, (Hants)3
- Batoids (Rajiformes, Torpediniformes, Myliobatiformes) from the Sülstorf Beds (Chattian, Late Oligocene) of Mecklenburg, northeastern Germany: a revision and description of three new species2
- Elasmobranch teeth (Vertebrata, Pisces) from the Dongen Formation (Eocene) in The Netherlands2
- Deep-Water Chondrichthyans from the Early Miocene of the Vienna Basin (Central Paratethys, Slovakia)1
- A Middle–Late Eocene vertebrate fauna (marine fish and mammals) from southwestern Morocco; preliminary report: age and palaeobiogeographical implications1
- Late Oligocene sharks and rays from the Chandler Bridge Formation, Dorchester County, South Carolina, USA1
- Contribution of Eocene sharks and rays from southern France to the history of deep-sea selachians1
- Structure and ichthyofauna of the upper part of Eocene sediments in Belogorsk area (Crimea)1
Each of the 46 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 13 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3
Picture of Gymnura
- Source
- Wikipedia/Commons
- Credit
- Tomas Willems & Hans Hillewaert
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Credit
- AMNH Mammal Collections
- Licence
- not stated by the source
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Echinosorex gymnura gymnura · AMNH:Mammals:M-216120
3D model
- Source
- Sketchfab
- Licence
- CC0 Public Domain
- Commercial use
- the licence allows it
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itGymnuridae through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Van Hasselt 1823