Hesperotestudo
Hesperotestudo is an extinct genus of tortoise native to North and Central America from the Early Miocene to the Late Pleistocene. Species of Hesperotestudo varied widely in size, with a large undescribed specimen from the Late Pleistocene of El Salvador reaching 150 cm (4.9 ft) in carapace length, larger than that of extant giant tortoises. Historically considered a subgenus of Geochelone, it is now considered to be distantly related to that genus. Its relationships with other tortoises are uncertain. The exposed areas of the bodies of Hesperotestudo species were extensively covered with large dermal ossicles, which in life were covered in keratin. It has been suggested that species of Hesperotestudo were relatively tolerant of cold weather. Hesperotestudo became extinct at the end of the Pleistocene roughly co-incident with the arrival of the first humans in North America. There is apparently a site in Florida where one individual may have been killed that some suggested were evidence of butchering, although others suggested that the turtle was neither cooked nor does a ledge that was found near it date at the same time as it.

A Cynical Idealist · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- XerobatinaePBDB
- Diet
- herbivore, herbivorePBDB, for Testudinidae
- Life habit
- ground dwellingPBDB, for Testudinidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Testudinidae
Time the record covers
At least 39 million years on record.
The record covers at least 39.7 Mya to 129 ka. No occurrence read is dated outside 45.9 Mya to 11.7 ka.
116 occurrences of Hesperotestudo on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
116 occurrences- Eocene2%
- Oligocene2%
- Miocene42%
- Pliocene14%
- Pleistocene41%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
35 formations
- Valentine Formation · Ogallala Group9
- Mehrten Formation6
- Ogallala Formation4
- Calvert Formation · Chesapeake Group4
- Bermont Formation3
- Rexroad Formation3
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The named rock units Hesperotestudo has been recovered from, largest first. Drawn where the rock is now, not where it formed. 11 further formations hold fewer occurrences each and are not listed. 51 occurrences have no formation on record, which is common for older literature.
Hesperotestudo and its relatives
MioceneWhere Hesperotestudo and its relatives were recorded in the Miocene.
Found in the same rock
1,732 taxa- Equus26 sites
- Lepisosteus20 sites
- Odocoileus virginianus20 sites
- Alligator mississippiensis19 sites
- Rana19 sites
- Terrapene carolina16 sites
- Thamnophis16 sites
- Dasypus bellus15 sites
- Holmesina septentrionalis15 sites
- Emydidae14 sites
- Hemiauchenia macrocephala14 sites
- Mammut americanum14 sites
- Gopherus polyphemus13 sites
- Mammuthus columbi13 sites
- Procyon lotor13 sites
- Pseudemys13 sites
- Hemiauchenia12 sites
- Kinosternon12 sites
- Nerodia12 sites
- Trionychidae12 sites
- Chelydra serpentina11 sites
- Lynx rufus11 sites
- Palaeolama mirifica11 sites
- Paramylodon harlani11 sites
The Paleobiology Database records 107 sites for Hesperotestudo, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
85 references- A fossil giant tortoise from the Mehrten Formation of Northern California6
- The Lapara Creek Fauna: Early Clarendonian of south Texas, USA3
- Pleistocene (Rancholabrean) vertebrate faunas from coastal Georgia3
- Pleistocene amphibians and reptiles from the Leisey Shell Pit, Hillsborough County, Florida3
- A new species of Geochelone from the Pleistocene of Texas3
- Miocene and Pliocene Chelonians from the Calvert Cliffs and Nearby Regions2
- Geometric morphometrics and anatomical network analyses reveal ecospace partitioning among geoemydid turtles from the Uinta Formation, Utah2
- The geology and paleontology of Ashfall Fossil Beds, a late Miocene (Clarendonian) mass-death assemblage, Antelope County and adjacent Knox County, Nebraska, USA2
- Amphibians and nonmarine turtles from the Miocene Calvert Formation of Delaware, Maryland, and Virginia (USA)2
- A Late Pleistocene Hepetofauna from Clark Quarry, Glynn County, Georgia2
- Testudines2
- The Tucker Borrow Pit: Paleontology and Stratigraphy of a Plio-Pleistocene fossil site in Brevard County, Florida2
Each of the 116 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 73 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Hesperotestudo
- Source
- Wikipedia/Commons
- Credit
- A Cynical Idealist
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Hesperotestudo was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itXerobatinae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Williams 1950)