Macronaria
Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located. Fossil evidence suggests that macronarian dinosaurs lived from the Middle Jurassic (Bathonian) through the Late Cretaceous (Maastrichtian). Macronarians have been found globally, including discoveries in Argentina, the United States, Portugal, China, and Tanzania. Like other sauropods, they are known to have inhabited primarily terrestrial areas, and little evidence exists to suggest that they spent much time in coastal environments. Macronarians are diagnosed through their distinct characters on their skulls, as well as appendicular and vertebral characters. Macronaria is composed of several subclades and families notably including Camarasauridae and Titanosauriformes, among several others. Titanosauriforms are particularly well known for being some of the largest terrestrial animals to ever exist.

User:Dhoefgen James St. John User:Adrian Grycuk User:Ghedoghedo User:SSR2000 User:Tiouraren · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- NeosauropodaPBDB
- Diet
- herbivorePBDB, for Sauropoda
- Life habit
- ground dwelling, gregariousPBDB, for Sauropoda
- Motility
- actively mobilePBDB, for Sauropoda
- Environment
- terrestrialPBDB, for Sauropoda
Time the record covers
At least 102 million years on record.
The record covers at least 174.7 Mya to 72.2 Mya. No occurrence read is dated outside 201.4 Mya to 66 Mya.
1,359 occurrences of Macronaria on record, most of them in the Late Cretaceous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
1,359 occurrences- Early Jurassicunder 1%
- Middle Jurassicunder 1%
- Late Jurassic20%
- Early Cretaceous19%
- Late Cretaceous60%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
276 formations
- Morrison Formation158
- Vitakri Formation · Fort Munro Group83
- Tendaguru Formation48
- Lameta Formation35
- Allen Formation · Malargüe Group33
- Winton Formation · Rolling Downs Group31
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The named rock units Macronaria has been recovered from, largest first. Drawn where the rock is now, not where it formed. 252 further formations hold fewer occurrences each and are not listed. 107 occurrences have no formation on record, which is common for older literature.
Macronaria and its relatives
Late CretaceousWhere Macronaria and its relatives were recorded in the Late Cretaceous.
Found in the same rock
945 taxa- Theropoda48 sites
- Sauropoda35 sites
- Giraffatitan brancai34 sites
- Titanosauridae33 sites
- Camarasaurus27 sites
- Titanosauria23 sites
- Testudines20 sites
- Dinosauria19 sites
- Crocodylia18 sites
- Titanosauriformes16 sites
- Allosaurus15 sites
- Diplodocus15 sites
- Dromaeosauridae15 sites
- Ornithopoda15 sites
- Allosaurus fragilis14 sites
- Stegosaurus14 sites
- Anura12 sites
- Abelisauridae11 sites
- Apatosaurus11 sites
- Coelurosauria11 sites
- Diplodocinae11 sites
- Alamosaurus sanjuanensis10 sites
- Kentrosaurus aethiopicus10 sites
- Lepidotes10 sites
The Paleobiology Database records 1,116 sites for Macronaria, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
733 references- New Balochisaurus (Balochisauridae, Titanosauria, Sauropoda) and Vitakridrinda (Theropoda) remains From Pakistan28
- Paleoecological analysis of the vertebrate fauna of the Morrison Formation (Upper Jurassic), Rocky Mountain region, U.S.A.24
- Die Gliedmaszen und Gliedmaszengürtel der Sauropoden der Tendaguru-Schichten [The limbs and limb girdles of the sauropods of the Tendaguru Beds]23
- Revision of South American titanosaurid dinosaurs: palaeobiological, palaeobiogeographical and phylogenetic aspects21
- Reappraisal of sauropod dinosaur diversity in the Upper Cretaceous Winton Formation of Queensland, Australia, through 3D digitisation and description of new specimens16
- Biodiversity of saurischian dinosaurs from the latest Cretaceous Park of Pakistan16
- Los vertebrados mesozoicos del Uruguay y sus relaciones con los de áreas vecinas [The Mesozoic vertebrates of Uruguay and their relationships with those of neighboring areas]16
- Review of Cretaceous sauropod dinosaurs from Central Asia15
- European titanosaurids15
- The Late Cretaceous dinosaur track record of Bolivia – review and perspective14
- Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriforms14
- Biostratigraphy of dinosaurs in the Upper Jurassic Morrison Formation of the Western Interior, U.S.A.13
Each of the 1,359 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 721 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Macronaria
- Source
- Wikipedia/Commons
- Credit
- User:Dhoefgen James St. John User:Adrian Grycuk User:Ghedoghedo User:SSR2000 User:Tiouraren
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Macronaria”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itNeosauropoda through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Sereno 1997