Phalacrocoracidae
Phalacrocoracidae is a family of approximately 40 species of aquatic birds found worldwide and commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the International Ornithologists' Union (IOU) adopted a consensus taxonomy of seven genera.

JJ Harrison ( https://www.jjharrison.com.au/ ) · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- cormorantPBDB
- Sits inside
- SuliformesPBDB
- Diet
- carnivorePBDB, for Coelurosauria
- Life habit
- ground dwelling, solitaryPBDB, for Theropoda
- Motility
- actively mobilePBDB, for Theropoda
- Environment
- terrestrialPBDB, for Theropoda
Time the record covers
At least 65 million years on record.
The record covers at least 66 Mya to 11.7 ka. No occurrence read is dated outside 72.2 Mya to 0 Mya.
327 occurrences of Phalacrocoracidae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
327 occurrences- Late Cretaceousunder 1%
- Eoceneunder 1%
- Oligocene5%
- Miocene13%
- Pliocene10%
- Pleistocene37%
- Holocene34%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
42 formations
- San Diego Formation13
- Hillgrove Formation9
- Pisco Formation8
- Namba Formation7
- Peace River Formation · Hawthorn Group5
- Adu Asa Formation5
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The named rock units Phalacrocoracidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 18 further formations hold fewer occurrences each and are not listed. 224 occurrences have no formation on record, which is common for older literature.
Phalacrocoracidae and its relatives
PleistoceneWhere Phalacrocoracidae and its relatives were recorded in the Pleistocene.
Found in the same rock
3,137 taxa- Phalacrocorax61 sites
- Phalacrocorax auritus38 sites
- Larus32 sites
- Phalacrocorax carbo29 sites
- Anas27 sites
- Phalacrocorax penicillatus27 sites
- Anas platyrhynchos25 sites
- Ardenna grisea25 sites
- Branta canadensis25 sites
- Anatinae23 sites
- Ardea herodias22 sites
- Equus22 sites
- Tyto alba22 sites
- Uria aalge22 sites
- Anserinae21 sites
- Phalacrocorax pelagicus21 sites
- Phoebastria albatrus21 sites
- Corvus corax20 sites
- Buteo jamaicensis19 sites
- Haliaeetus leucocephalus19 sites
- Anser caerulescens18 sites
- Buteo18 sites
- Numenius americanus18 sites
- Podiceps nigricollis18 sites
The Paleobiology Database records 219 sites for Phalacrocoracidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
145 references- Prehistoric human impacts on California birds: evidence from the Emeryville shellmoumd avifauna54
- A remarkable fossil and archaeological avifauna from Marfells Beach, Lake Grassmere, South Island, New Zealand19
- Fossil birds of the San Diego Formation, Late Pliocene, Blancan, San Diego County, California11
- Seasonal exploitation of seabirds by Late Holocene coastal foragers: Analysis of modern and archaeological data from Western Cape, South Africa10
- Late-Pleistocene avifaunas from Cape Wanbrow, Otago, South Island, New Zealand9
- Seabirds and Late Pleistocene marine environments in the Northeast Atlantic and the Mediterranean8
- Descriptions and phylogenetic relationships of a new genus and two new species of Oligo-Miocene cormorants (Aves: Phalacrocoracidae) from Australia7
- A catalogue of British Pleistocene birds identified by Colin JO Harrison and stored in the Natural History Museum, London, (Department of Palaeontology)7
- Distribution and provenance of fossil avifauna on San Miguel Island7
- Fossil birds from the Late Miocene of Chad and Ethiopia and zooge- ographical implications5
- A late Pleistocene avifauna from San Miguel Island, California5
- Terminal Pleistocene–Holocene Avifauna of San Miguel and Santa Rosa Islands: Identifications of Previously Unidentified Avian Remains Recovered from Fossil Sites and Prehistoric Cave Deposits4
Each of the 327 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 133 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Phalacrocoracidae
- Source
- Wikipedia/Commons
- Credit
- JJ Harrison ( https://www.jjharrison.com.au/ )
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Phalacrocoracidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Cormorant”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itSuliformes through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Reichenbach 1850