Cephalocarida
The Cephalocarida are a class in the subphylum Crustacea comprising only 12 species. Both the nauplii and the adults are benthic. They were discovered in 1955 by Howard L. Sanders, and are commonly referred to as horseshoe shrimp. They have been grouped together with the Remipedia in the Xenocarida. Although a second family, Lightiellidae, is sometimes used, all cephalocaridans are generally considered to belong in just one family: Hutchinsoniellidae. Fossil records of cephalocaridans have been found in the Ordovician Castle Bank site.

Eric A. Lazo-Wasem · CC0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- AllotriocaridaPBDB
- Diet
- detritivorePBDB
- Life habit
- low-level epifaunalPBDB
- Motility
- slow-movingPBDB
- Vision
- blindPBDB
Time the record covers
At least 24 million years on record.
The record covers at least 493.6 Mya to 469.4 Mya. No occurrence read is dated outside 497 Mya to 458.2 Mya.
2 occurrences of Cephalocarida on record, most of them in the Cambrian. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
2 occurrences- Cambrian50%
- Ordovician50%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
2 formations
- Alum Shale Formation1
- Gilwern Volcanic Formation · Builth Volcanic Group1
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The named rock units Cephalocarida has been recovered from, largest first. Drawn where the rock is now, not where it formed.
Cephalocarida and its relatives
CambrianWhere Cephalocarida and its relatives were recorded in the Cambrian.
Found in the same rock
47 taxa- Acrotretida1 site
- Apatobolus1 site
- Archaeopriapulida1 site
- Arthropoda1 site
- Ascospongiella1 site
- Asterozoa1 site
- Bredocaris admirabilis1 site
- Chaetetida1 site
- Chitinozoa1 site
- Cirripedia1 site
- Conodonta1 site
- Conulariida1 site
- Cyathophycus1 site
- Dala1 site
- Dendrocystites1 site
- Dictyonema1 site
- Didymograptus murchisoni1 site
- Diplograptus1 site
- Falites1 site
- Fenestellida1 site
- Filisphaeridium1 site
- Graptolithina1 site
- Herpetogaster1 site
- Hesslandona1 site
The Paleobiology Database records 2 sites for Cephalocarida, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
2 references- A Middle Ordovician Burgess Shale-type fauna from Castle Bank, Wales (UK)1
- Crustacea with preserved soft parts from the Upper Cambrian of Sweden1
Each of the 2 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Each entry links to doi.org. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Cephalocarida
- Source
- Wikipedia/Commons
- Credit
- Eric A. Lazo-Wasem
- Licence
- CC0
- Commercial use
- the licence allows it
No 3D scan of Cephalocarida was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAllotriocarida through timeFormations of the CambrianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Sanders 1955