Graptolithina
Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian through the Lower Carboniferous (Mississippian). A possible early graptolite, Chaunograptus, is known from the Middle Cambrian. Recent analyses have favored the idea that the living pterobranch Rhabdopleura represents an extant graptolite which diverged from the rest of the group in the Cambrian.

Grezmagro · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- graptolitePBDB
- Sits inside
- PterobranchiaPBDB
- Diet
- suspension feederPBDB
- Life habit
- planktonicPBDB
- Motility
- passively mobilePBDB
- Vision
- blindPBDB
Time the record covers
At least 157 million years on record.
The record covers at least 504.5 Mya to 346.7 Mya. No occurrence read is dated outside 508.2 Mya to 339.9 Mya.
8,119 occurrences of Graptolithina on record, most of them in the Ordovician. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
8,119 occurrences- Cambrianunder 1%
- Ordovician66%
- Silurian29%
- Devonian4%
- Carboniferousunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
678 formations
- Phi Kappa Formation · Valmy Group357
- Vinini Formation315
- Wufeng Formation209
- Lower Road River Formation · Road River Group201
- Aorangi Mine Formation199
- Road River Formation151
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The named rock units Graptolithina has been recovered from, largest first. Drawn where the rock is now, not where it formed. 654 further formations hold fewer occurrences each and are not listed. 1,398 occurrences have no formation on record, which is common for older literature.
Graptolithina and its relatives
OrdovicianWhere Graptolithina and its relatives were recorded in the Ordovician.
Found in the same rock
2,073 taxa- Climacograptus72 sites
- Didymograptus60 sites
- Glyptograptus47 sites
- Orthograptus42 sites
- Dicellograptus41 sites
- Remopleurides26 sites
- Diplograptus25 sites
- Amplexograptus24 sites
- Lonchodomas23 sites
- Megalaspides23 sites
- Tetragraptus23 sites
- Lingula22 sites
- Phyllograptus22 sites
- Dictyonema21 sites
- Lingulella21 sites
- Ampyx20 sites
- Ctenodonta19 sites
- Hyolithes19 sites
- Onniella19 sites
- Illaenus18 sites
- Sowerbyella18 sites
- Leptaena17 sites
- Trinodus16 sites
- Dicranograptus15 sites
The Paleobiology Database records at least 603 sites for Graptolithina, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
604 references- Uppermost Ordovician (upper Katian-Hirnantian) graptolites of north-central Nevada, USA382
- Ordovician and Silurian graptolite succession in the Trail Creek area, central Idaho-a graptolite zone reference section344
- Correlation of the Silurian rocks of China340
- Ordovician Sedimentary Basin and Paleobiotas of the Gorny Altai327
- Ordovician Geology and Graptolite faunas of the Aorangi Mine area, North-west Nelson, New Zealand230
- Correlation of the African Silurian rocks214
- Graptolites of Cobb River-Mount Arthur Area, North-West Nelson, New Zealand178
- Stratigraphic Summary of the Southwestern Region of China168
- Regional stratigraphic table of Eastern China, Zhejiang Province158
- Ordovician, Silurian, and Devonian biostratigraphy of east-central Alaska157
- Regional stratigraphic tables of the north western China region: Volume of NinXia Province148
- Llandoverian and Wenlockian graptolites from Bornholm145
Each of the 8,119 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 592 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Graptolithina
- Source
- Wikipedia/Commons
- Credit
- Grezmagro
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Graptolithina was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Graptolite”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPterobranchia through timeFormations of the OrdovicianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Bronn 1849)