Chelicerata
The subphylum Chelicerata constitutes one of the two major living subdivisions of the phylum Arthropoda, alongside Mandibulata. Chelicerates include the sea spiders, horseshoe crabs, and arachnids, as well as a number of extinct lineages, such as the eurypterids and chasmataspidids.

Ken-ichi Ueda Hans Hillewaert Luis Fernández García Lucarelli Bernard DUPONT Kurt Kulac · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- DeuteropodaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
Time the record covers
At least 515 million years on record.
The record covers at least 516 Mya to 11.7 ka. No occurrence read is dated outside 538.8 Mya to 0 Mya.
4,901 occurrences of Chelicerata on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
4,901 occurrences- Cambrianunder 1%
- Ordovicianunder 1%
- Silurian8%
- Devonian6%
- Carboniferous13%
- Permianunder 1%
- Early Triassicunder 1%
- Middle Triassicunder 1%
- Late Triassicunder 1%
- Early Jurassicunder 1%
- Middle Jurassicunder 1%
- Late Jurassicunder 1%
- Early Cretaceous3%
- Late Cretaceous14%
- Paleoceneunder 1%
- Eocene38%
- Oligoceneunder 1%
- Miocene10%
- Plioceneunder 1%
- Pleistoceneunder 1%
- Holocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
464 formations
- Carbondale Formation · Kewanee Group95
- Florissant Formation57
- Kladno Formation46
- Crato Formation · Santana Group34
- Fiddlers Green Formation · Bertie Group32
- Niveau du gypse d'Aix Formation26
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The named rock units Chelicerata has been recovered from, largest first. Drawn where the rock is now, not where it formed. 440 further formations hold fewer occurrences each and are not listed. 3,332 occurrences have no formation on record, which is common for older literature.
Chelicerata and its relatives
EoceneWhere Chelicerata and its relatives were recorded in the Eocene.
Found in the same rock
3,608 taxa- Bivalvia37 sites
- Pterygotus31 sites
- Gastropoda26 sites
- Orbiculoidea26 sites
- Ostracoda26 sites
- Howellella25 sites
- Eurypterida24 sites
- Lingula24 sites
- Leperditia22 sites
- Crinoidea21 sites
- Cephalopoda20 sites
- Camarotoechia18 sites
- Nuculites18 sites
- Phyllocarida17 sites
- Plantae16 sites
- Trilobita16 sites
- Carcinosoma15 sites
- Brachiopoda14 sites
- Bryozoa14 sites
- Erettopterus14 sites
- Erieopterus14 sites
- Tentaculites14 sites
- Hughmilleria13 sites
- Conularia12 sites
The Paleobiology Database records at least 483 sites for Chelicerata, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
1,043 references- Habitat of Llandoverian-Lochkovian eurypterids246
- Die fossilen Spinnen im dominikanischen Bernstein186
- On extant and fossil (Eocene) European comb-footed spiders (Araneae: Theridiidae), with notes on their subfamilies, and with descriptions of new taxa174
- Footnotes and taxonomic names, in Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt136
- Die im Bernstein befindlichen Crustaceen, Myriapoden, Arachniden und Apteren der Vorwelt124
- On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida: Araneae)97
- New and rare fossil spiders (Araneae) in mid Cretaceous amber from Myanmar (Burma), including the description of new extinct families of the suborders Mesothelae and Opisthothelae, as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae88
- A study of amber spiders88
- Amber Spiders in European Collections84
- Descriptions of new fossil spiders (Arachnida: Araneae) in Late (Mid) Cretaceous Burmese amber with focus on the superfamilies Palpimanoidea and Deinopoidea and members of the RTA-clade, as well as remarks on palaeobehaviour, palaeofauna, taxonomy and phylogenetics70
- The fossil spiders of the family Anapidae s. l. (Araneae) in Baltic, Dominican and Mexican amber and their extant relatives, with the description of a new subfamily Comarominae68
- A restudy of the fossil Scorpionida of the world57
Each of the 4,901 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 1,031 further references are not listed. Each entry links to the Paleobiology Database's own record for the work.
Media and terms
Media content2
Picture of Chelicerata
- Source
- Wikipedia/Commons
- Credit
- Ken-ichi Ueda Hans Hillewaert Luis Fernández García Lucarelli Bernard DUPONT Kurt Kulac
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
3D model
- Source
- Sketchfab
- Licence
- CC Attribution-ShareAlike
- Commercial use
- cannot be read from the licence given — check the record
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Chelicerata”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itDeuteropoda through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Heymons 1901