Gymnuridae
The butterfly rays are the rays forming the genus Gymnura and the family Gymnuridae. They are found in warm oceans worldwide, and occasionally in estuaries.

Tomas Willems & Hans Hillewaert · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- MyliobatoideiPBDB
- Diet
- carnivorePBDB, for Myliobatiformes
- Life habit
- nektobenthicPBDB, for Myliobatiformes
- Motility
- actively mobilePBDB, for Myliobatiformes
Time the record covers
At least 88 million years on record.
The record covers at least 93.9 Mya to 5.333 Mya. No occurrence read is dated outside 100.5 Mya to 3.6 Mya.
109 occurrences of Gymnuridae on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
109 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
43 formations
- Birket Qarun Formation9
- Matam Formation8
- Gehannam Formation5
- Couche II Formation5
- Qasr el-Sagha Formation4
- Marnes de Donzacq Formation4
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The named rock units Gymnuridae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 19 further formations hold fewer occurrences each and are not listed. 15 occurrences have no formation on record, which is common for older literature.
Gymnuridae and its relatives
EoceneWhere Gymnuridae and its relatives were recorded in the Eocene.
Found in the same rock
1,261 taxa- Dasyatis41 sites
- Gymnura30 sites
- Myliobatis26 sites
- Rhinobatos26 sites
- Coupatezia25 sites
- Scyliorhinus21 sites
- Rhizoprionodon20 sites
- Physogaleus secundus18 sites
- Jacquhermania duponti17 sites
- Striatolamia macrota17 sites
- Burnhamia daviesi16 sites
- Rhinoptera16 sites
- Heterodontus vincenti15 sites
- Anoxypristis14 sites
- Brachycarcharias lerichei14 sites
- Hypotodus verticalis14 sites
- Odontaspis winkleri14 sites
- Rhynchobatus14 sites
- Abdounia13 sites
- Ouledia13 sites
- Premontreia (Oxyscyllium) gilberti13 sites
- Pristis13 sites
- Coupatezia woutersi12 sites
- Galeorhinus12 sites
The Paleobiology Database records 80 sites for Gymnuridae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
48 references- Shark and ray faunas in the Middle and Late Eocene of the Fayum Area, Egypt14
- Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia9
- Les Orectolobiformes, Carcharhiniformes et Myliobatiformes (Elasmobranchii, Neoselachii) des bassins à phosphate du Maroc (Maastrichtien-Lutétien basal). Systématique, biostratigraphie, évolution et dynamique des faunes9
- The selachian fauna (sharks and rays) of the phosphate series of Ndendouri-Ouali Diala (Matam, Western Senegal): Dating and paleoenvironmental interests8
- Nouvelle faunes de sélaciens (Elasmobranchii, Neoselachii) de l'Eocène moyen des Landes (Sud-Ouest, France). Implication dans la connaissance des communautés de sélaciens d'eau profondes7
- Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths3
- Sélaciens du Miocène terminal du bassin d'Alvalade (Portugal) Essai de synthèse3
- Un noveau genre de sélacien (Batomorphii, Myliobatiformes) de l'Yprésien des Ouled Abdoun, Maroc)3
- The Selsey Division (Bracklesham Group) at Lee-on-the-Solent, Gosport, (Hants)3
- A selachian fauna from the middle Eocene (Lutetian, Lisbon Formation) of Andalusia, Covington County, Alabama, USA2
- Batoids (Rajiformes, Torpediniformes, Myliobatiformes) from the Sülstorf Beds (Chattian, Late Oligocene) of Mecklenburg, northeastern Germany: a revision and description of three new species2
- A Middle–Late Eocene vertebrate fauna (marine fish and mammals) from southwestern Morocco; preliminary report: age and palaeobiogeographical implications2
Each of the 109 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 36 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Gymnuridae
- Source
- Wikipedia/Commons
- Credit
- Tomas Willems & Hans Hillewaert
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMyliobatoidei through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Fowler 1934