Hemichordata
Hemichordata is a phylum which consists of triploblastic, eucoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include two main classes: Enteropneusta, and Pterobranchia. A third class, Planctosphaeroidea, is known only from the larva of a single species, Planctosphaera pelagica. The class Graptolithina, formerly considered extinct, is now placed within the pterobranchs, represented by a single living genus Rhabdopleura.

Gustav Paulay · CC0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- acorn wormPBDB
- Sits inside
- AmbulacrariaPBDB
Time the record covers
At least 326 million years on record.
The record covers at least 525.5 Mya to 199.5 Mya. No occurrence read is dated outside 538.8 Mya to 192.9 Mya.
8,194 occurrences of Hemichordata on record, most of them in the Ordovician. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
8,194 occurrences- Cambrian1%
- Ordovician66%
- Silurian29%
- Devonian4%
- Carboniferousunder 1%
- Early Jurassicunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
694 formations
- Phi Kappa Formation · Valmy Group357
- Vinini Formation315
- Wufeng Formation209
- Lower Road River Formation · Road River Group201
- Aorangi Mine Formation199
- Road River Formation151
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The named rock units Hemichordata has been recovered from, largest first. Drawn where the rock is now, not where it formed. 670 further formations hold fewer occurrences each and are not listed. 1,399 occurrences have no formation on record, which is common for older literature.
Hemichordata and its relatives
OrdovicianWhere Hemichordata and its relatives were recorded in the Ordovician.
Found in the same rock
2,074 taxa- Climacograptus72 sites
- Didymograptus60 sites
- Glyptograptus47 sites
- Orthograptus42 sites
- Dicellograptus41 sites
- Remopleurides26 sites
- Diplograptus25 sites
- Amplexograptus24 sites
- Lonchodomas23 sites
- Megalaspides23 sites
- Tetragraptus23 sites
- Lingula22 sites
- Phyllograptus22 sites
- Dictyonema21 sites
- Lingulella21 sites
- Ampyx20 sites
- Ctenodonta19 sites
- Hyolithes19 sites
- Onniella19 sites
- Illaenus18 sites
- Sowerbyella18 sites
- Leptaena17 sites
- Trinodus16 sites
- Dicranograptus15 sites
The Paleobiology Database records at least 602 sites for Hemichordata, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
626 references- Uppermost Ordovician (upper Katian-Hirnantian) graptolites of north-central Nevada, USA382
- Ordovician and Silurian graptolite succession in the Trail Creek area, central Idaho-a graptolite zone reference section344
- Correlation of the Silurian rocks of China340
- Ordovician Sedimentary Basin and Paleobiotas of the Gorny Altai327
- Ordovician Geology and Graptolite faunas of the Aorangi Mine area, North-west Nelson, New Zealand230
- Correlation of the African Silurian rocks214
- Graptolites of Cobb River-Mount Arthur Area, North-West Nelson, New Zealand178
- Stratigraphic Summary of the Southwestern Region of China168
- Regional stratigraphic table of Eastern China, Zhejiang Province158
- Ordovician, Silurian, and Devonian biostratigraphy of east-central Alaska157
- Regional stratigraphic tables of the north western China region: Volume of NinXia Province148
- Llandoverian and Wenlockian graptolites from Bornholm145
Each of the 8,194 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 614 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Hemichordata
- Source
- Wikipedia/Commons
- Credit
- Gustav Paulay
- Licence
- CC0
- Commercial use
- the licence allows it
3D model
- Source
- Sketchfab
- Licence
- CC0 Public Domain
- Commercial use
- the licence allows it
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itAmbulacraria through timeFormations of the OrdovicianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Bateson 1885