Holocephali
Searched as Holocephali indet.. The Paleobiology Database treats Holocephali as the accepted name.
Holocephali is a subclass of cartilaginous fish. The only living holocephalans are the three families of chimaeras, but the group also includes many extinct members and was more diverse during the Paleozoic and Mesozoic eras. The earliest known fossils of holocephalans date to the Middle Devonian, and the subclass likely reached its peak diversity during the following Carboniferous Period. Molecular clock studies suggest that holocephalans diverged from their closest relatives, elasmobranchs such as sharks and rays, during the Early Devonian or the Silurian Period.

Montanoceratops · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- chimaeraPBDB
- Sits inside
- ChondrichthyesPBDB
- Diet
- carnivorePBDB
- Life habit
- nektobenthicPBDB
- Motility
- actively mobilePBDB
Time the record covers
At least 292 million years on record.
The record covers at least 303.7 Mya to 11.63 Mya. No occurrence read is dated outside 307 Mya to 7.246 Mya.
11 occurrences of Holocephali on record, most of them in the Late Triassic. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
11 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
8 formations
- Parana Formation2
- Sables de Varennes Formation1
- Khodzhakul Formation1
- Grès de Infralias Formation1
- Mattoon Formation1
- Woburn Sands Formation1
Loading coastlines…
The named rock units Holocephali has been recovered from, largest first. Drawn where the rock is now, not where it formed. 2 occurrences have no formation on record, which is common for older literature.
Holocephali and its relatives
Late TriassicWhere Holocephali and its relatives were recorded in the Late Triassic.
Found in the same rock
2,709 taxa- Ischyrhiza mira15 sites
- Petalodus15 sites
- Teleostei13 sites
- Crocodylia12 sites
- Glikmanius occidentalis12 sites
- Cretalamna appendiculata11 sites
- Anomoeodus phaseolus10 sites
- Ischyodus bifurcatus10 sites
- Archaeolamna kopingensis9 sites
- Bivalvia9 sites
- Scapanorhynchus texanus9 sites
- Composita8 sites
- Crinoidea8 sites
- Enchodus8 sites
- Enchodus ferox8 sites
- Ischyodus8 sites
- Orbiculoidea8 sites
- Osteichthyes8 sites
- Paralbula casei8 sites
- Scyliorhinus8 sites
- Squalicorax kaupi8 sites
- Squalicorax pristodontus8 sites
- Squatina8 sites
- Testudines8 sites
The Paleobiology Database records 637 sites for Holocephali, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
9 references- Microvertebrates from the Lower Greensand Group (Lower Cretaceous) of Clophill, Bedfordshire, UK, and Nutfield, Surrey, UK2
- L'ichthyofaune du Pliensbachian (Jurassique Inférieur) de Lorraine et des Ardennes (France): premiers résultats2
- A taxonomic and paleoecological review of the Rhaetian chondrichthyan fauna of Bonenburg (NRW) Germany1
- Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic1
- Fishes and tetrapods in the upper Pennsylvanian (Kasimovian) Cohn Coal Member of the Mattoon formation of Illinois, United States: systematics, paleoecology, and paleoenvironments1
- A sperm whale (Cetacea: Physeteroidea) from the Parana Formation (Late Miocene) of Entre Rios, Argentina. Environment and taphonomy1
- Une dent de Nodosauridé (Dinosauria, Ankylosauria) dans le Cénomanien de Touraine [A nodosaurid tooth (Dinosauria, Ankylosauria) from the Cenomanian of Touraine]1
- Evolution des faunes de vertébrés à la limite trias-jurassique: apports de la Lorraine [Evolution of vertebrate faunas at the Triassic-Jurassic boundary: contributions from Lorraine]1
- Cretaceous anurans from Central Asia1
Each of the 11 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Holocephali
- Source
- Wikipedia/Commons
- Credit
- Montanoceratops
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Holocephali was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Holocephali”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itChondrichthyes through timeFormations of the Late TriassicThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Bonaparte 1832)